Vol.:(0123456789) Archives of Virology (2022) 167:2857–2906 https://doi.org/10.1007/s00705-022-05546-z VIROLOGY DIVISION NEWS 2022 taxonomic update of phylum Negarnaviricota (Riboviria: Orthornavirae), including the large orders Bunyavirales and Mononegavirales Jens H. Kuhn1  · Scott Adkins2  · Sergey V. Alkhovsky (Aльxoвcкий Cepгeй Bлaдимиpoвич) 3  · Tatjana Avšič‑Županc4  · María A. Ayllón5,6  · Justin Bahl7  · Anne Balkema‑Buschmann8  · Matthew J. Ballinger9  · Martina Bandte10  · Martin Beer11  · Nicolas Bejerman12  · Éric Bergeron13  · Nadine Biedenkopf14  · Laurent Bigarré15  · Carol D. Blair16  · Kim R. Blasdell17  · Steven B. Bradfute18  · Thomas Briese19  · Paul A. Brown20  · Rémy Bruggmann21  · Ursula J. Buchholz22  · Michael J. Buchmeier23  · Alexander Bukreyev24,91  · Felicity Burt25  · Carmen Büttner10  · Charles H. Calisher26 · Thierry Candresse27  · Jeremy Carson28  · Inmaculada Casas29  · Kartik Chandran30  · Rémi N. Charrel31  · Yuya Chiaki  (千秋祐也) 32  · Anya Crane33  · Mark Crane34 · Laurent Dacheux35  · Elena Dal Bó36 · Juan Carlos de la Torre37  · Xavier de Lamballerie38 · William M. de Souza39  · Rik L. de Swart40  · Nolwenn M. Dheilly41  · Nicholas Di Paola42  · Francesco Di Serio43  · Ralf G. Dietzgen44  · Michele Digiaro45 · J. Felix Drexler46  · W. Paul Duprex47  · Ralf Dürrwald48  · Andrew J. Easton49  · Toufic Elbeaino45  · Koray Ergünay50,51,52,53  · Guozhong Feng (冯国忠) 54  · Claudette Feuvrier55 · Andrew E. Firth56  · Anthony R. Fooks57  · Pierre B. H. Formenty58  · Juliana Freitas‑Astúa59  · Selma Gago‑Zachert60  · María Laura García61  · Adolfo García‑Sastre62  · Aura R. Garrison63  · Scott E. Godwin28  · Jean‑Paul J. Gonzalez64  · Joëlle Goüy de Bellocq65  · Anthony Griffiths66  · Martin H. Groschup67  · Stephan Günther68  · John Hammond69  · Jussi Hepojoki70,71  · Melanie M. Hierweger72 · Seiji Hongō (本郷誠治) 73 · Masayuki Horie (堀江真行) 74,75  · Hidenori Horikawa (堀川英則) 76 · Holly R. Hughes77  · Adam J. Hume78,79  · Timothy H. Hyndman80  · Dàohóng Jiāng (姜道宏) 81  · Gilda B. Jonson82  · Sandra Junglen46  · Fujio Kadono  (上遠野冨士夫) 83  · David G. Karlin10  · Boris Klempa84  · Jonas Klingström85  · Michel C. Koch86  · Hideki Kondō (近藤秀樹) 87  · Eugene V. Koonin88 · Jarmila Krásová65 · Mart Krupovic89  · Kenji Kubota  (久保田健嗣) 90  · Ivan V. Kuzmin91  · Lies Laenen92,93  · Amy J. Lambert94  · Jiànróng Lǐ (李建荣) 95 · Jun‑Min Li (李俊敏) 96  · François Lieffrig97 · Igor S. Lukashevich98  · Dongsheng Luo99,100 · Piet Maes101  · Marco Marklewitz102  · Sergio H. Marshall103  · Shin‑Yi L. Marzano104  · John W. McCauley105  · Ali Mirazimi106 · Peter G. Mohr34  · Nick J. G. Moody34 · Yasuaki Morita (森田泰彰) 107 · Richard N. Morrison28  · Elke Mühlberger108  · Rayapati Naidu109  · Tomohide Natsuaki (夏秋知英) 110  · José A. Navarro111  · Yutaro Neriya (煉谷裕太朗) 112  · Sergey V. Netesov (Heтёcoв Cepгeй Bиктopoвич) 113  · Gabriele Neumann114  · Norbert Nowotny115,116  · Francisco M. Ochoa‑Corona117  · Gustavo Palacios118  · Laurane Pallandre119 · Vicente Pallás120  · Anna Papa (Άννα Παπά) 121  · Sofia Paraskevopoulou (Σοφία Παρασκευοπούλου) 122  · Colin R. Parrish123 · Alex Pauvolid‑Corrêa124  · Janusz T. Pawęska125  · Daniel R. Pérez126  · Florian Pfaff55 · Richard K. Plemper127  · Thomas S. Postler128  · Françoise Pozet55 · Sheli R. Radoshitzky129  · Pedro L. Ramos‑González130  · Marius Rehanek10  · Renato O. Resende131  · Carina A. Reyes132  · Víctor Romanowski133  · Dennis Rubbenstroth134  · Luisa Rubino43  · Artemis Rumbou (Άρτεμις Ρούμπου) 10  · Jonathan A. Runstadler135  · Melanie Rupp136 · Sead Sabanadzovic137  · Takahide Sasaya  (笹谷孝英) 138  · Heike Schmidt‑Posthaus139  · Martin Schwemmle140  · Torsten Seuberlich141  · Stephen R. Sharpe142  · Mang Shi (施莽) 143  · Manuela Sironi144  · Sophie Smither145  · Jin‑Won Song  (송진원) 146  · Kirsten M. Spann147  · Jessica R. Spengler148  · Mark D. Stenglein149  · Ayato Takada  (高田礼人) 150  · Robert B. Tesh91 · Jana Těšíková65,151 · Natalie J. Thornburg152  · Nicole D. Tischler153  · Yasuhiro Tomitaka (冨髙保弘) 154  · Keizō Tomonaga (朝長啓造) 155  · Noël Tordo156  · Kenta Tsunekawa157 · Handling Editor: Tim Skern. Extended author information available on the last page of the article 2858 J. H. Kuhn et al. 1 3 Massimo Turina158  · Ioannis E. Tzanetakis159  · Anna Maria Vaira158  · Bernadette van den Hoogen160  · Bert Vanmechelen161  · Nikos Vasilakis (Νίκος Βασιλάκης) 91  · Martin Verbeek162  · Susanne von Bargen10  · Jiro Wada (和田治郎) 163  · Victoria Wahl164  · Peter J. Walker165  · Anna E. Whitfield166  · John V. Williams167  · Yuri I. Wolf168  · Junki Yamasaki (山﨑淳紀) 169 · Hironobu Yanagisawa (栁澤広宣) 170  · Gongyin Ye (叶恭银) 171  · Yong‑Zhen Zhang (张永振) 172 · Arnfinn Lodden Økland173 Published online: 28 November 2022 This is a US Government work and not under copyright protection in the US; foreign copyright protection may apply 2022 Abstract In March 2022, following the annual International Committee on Taxonomy of Viruses (ICTV) ratification vote on newly proposed taxa, the phylum Negarnaviricota was amended and emended. The phylum was expanded by two new families (bunyaviral Discoviridae and Tulasviridae), 41 new genera, and 98 new species. Three hundred forty-nine species were renamed and/or moved. The accidentally misspelled names of seven species were corrected. This article presents the updated taxonomy of Negarnaviricota as now accepted by the ICTV. Introduction The phylum Negarnaviricota was established in 2019 by the International Committee on Taxonomy of Viruses (ICTV) for negative-sense RNA viruses that can be connected evo- lutionarily through the possession of virally encoded RNA- directed RNA polymerases (RdRps). The phylum includes two subphyla, Haploviricotina and Polyploviricotina, for negative-sense RNA viruses that encode RdRps with or without mRNA capping activity, respectively. The two sub- phyla include four classes (Chunqiuviricetes, Milnevirice- tes, Monjiviricetes, and Yunchangviricetes) and two classes (Ellioviricetes and Insthoviricetes), respectively [33, 59, 74]. The phylum was last amended/emended in 2021 [34]. Here, we present the changes that were proposed to the phylum via official taxonomic proposals (TaxoProps) in 2021 and accepted by the ICTV in March and May 2022 [23, 70]. These changes are now part of the official ICTV taxonomy [24]. Taxonomic changes in subphylum Haploviricotina Chunqiuviricetes The names of all species in the class were changed to binomi- als as required by the recently amended International Code of Virus Classification and Nomenclature (ICVCN) [25, 69, 81] (TaxoProp 2021.018M.R.Negarnaviricota_sprename). Milneviricetes The names of all species in the class were changed to bino- mials as required by the recently amended ICVCN [25, 69, 81] (TaxoProp 2021.014P.R.Aspiviridae_rename). Monjiviricetes Family Aliusviridae The species name Olluvirus shayangense was corrected to Ollusvirus shayangense (TaxoProp 2021.042M.R.Corrections_Riboviria_Duplodnaviria). Family Artoviridae The names of all species in the family were changed to binomials as required by the recently amended ICVCN [25, 69, 81] (TaxoProp 2021.009M.R.Artoviridae_sprename). Genus Peropuvirus was expanded by one new species, Peropuvirus melongenae, for Solanum melongena rhabdo- like virus (SmRLV) identified in a metagenomic study of eggplant (Solanum melongena L.) leaf tissue sampled in Zhènjiāng (镇江市), Jiāngsū Province (江苏省), China [78] (TaxoProp 2021.029M.R.Peropuvirus_1nsp). Family Bornaviridae The names of all species in the family were changed to binomials as required by the recently amended ICVCN [25, 69, 81] (2021.010M.R.Bornaviridae_sprename). Genus Orthobornavirus was expanded by one new spe- cies, Orthobornavirus caenophidiae, for Caribbean water- snake bornavirus (CWBV) and Mexican black-tailed rat- tlesnake bornavirus (MRBV) identified in archived raw transcriptomic read data of a Caribbean watersnake (colubrid Tretanorhinus variabilis A.M.C. Duméril, Bibron & A.H.A. 2859Taxonomic update of phylum Negarnaviricota 1 3 Duméril, 1854) and a Mexican black-tailed rattlesnake (vip- erid Crotalus molossus nigrescens Gloyd, 1936), respectively [49] (TaxoProp 2021.021M.R.Orthobornavirus_1nsp). in Hángzhōu (杭州市), Zhèjiāng Province (浙江省), China [72]; • genus Copasivirus was created to include two new spe- cies: o Copasivirus manlyvaleense for Jimsystermes virus (JIMV) discovered by HTS in termites (termitid Occasitermes sp.) sampled in Manly Vale, New South Wales, Australia [35]; and o Copasivirus ivindoense for isopteran arli-related virus OKIAV103 (IARV-103) discovered by HTS in termites (rhinotermitid Coptotermes sp.) sampled in Ivindo National Park (Parc national d'Ivindo), Ogooué-Ivindo Province, Gabon [27]; • genus Cybitervirus was created to include new species Cybitervirus niederense for coleopteran arli-related virus OKIAV107 (CARV-107) discovered by HTS in beetles (dytiscid Cybister lateralimarginalis (De Geer, 1774)) sampled in Lüchow-Dannenberg, Lower Saxony (Nied- ersachsen), Germany [27]; • genus Phelinovirus was created to include new species Phelinovirus aphidis for hymenopteran arli-related virus OKIAV99 (HARV-99) discovered by HTS in parasi- toid wasps (aphelinid Aphelinus abdominalis (Dalman, 1820)) in laboratory culture samples of unknown geo- graphical origin [27]; • genus Rivapovirus was created to include new species Rivapovirus aleyrodidae for hemipteran arli-related virus OKIAV94 (HARV-94) discovered by HTS in greenhouse whiteflies (aleyrodid Trialeurodes vaporariorum West- wood, 1856) of commercial laboratory cultures sampled in Brandenburg, Germany [27]; • genus Stylovirus was created to include new species Stylovirus niederense for strepsipteran arli-related virus OKIAV104 (SARV-104) discovered by HTS in endo- parasitic insects (stylopid Stylops melittae Kirby, 1802) sampled in Osnabrück, Lower Saxony (Niedersachsen), Germany [27]; • genus Supelovirus was created to include the new species Supelovirus thailandense for blattodean arli-related virus OKIAV102 (BARV-102) discovered by HTS in brown- banded cockroaches (ectobiid Supella longipalpa Fab- ricius, 1798) of laboratory culture samples originating from Thailand [27]; • genus Synelinevirus was created to include two new spe- cies: o Synelinevirus paranaense for Linepithema humile rhabdo-like virus 1 (LhuRLV-1) 1 discovered by HTS in Argentine ants (dolichoderine Linepithema Family Filoviridae Family Filoviridae was expanded by two genera: • genus Oblavirus was established to include one new species, Oblavirus percae, for Oberland virus (OBLV) discovered by high-throughput sequencing (HTS) in European perch (percid Perca fluviatilis Linnaeus, 1758) sampled in Switzerland [20] (TaxoProp 2021.011M.R.Fi loviridae_1ngen_3nsp); and • genus Tapjovirus was established to include one new species, Tapjovirus bothropis, for Tapajós virus (TAPV) discovered in sequence read archive (SRA) data obtained from a common lancehead (viperid Bothrops atrox (Linnaeus, 1758)) sampled in Brazil [21] (TaxoProp 2021.005M.R.Filoviridae_1ngen_1nsp). Genus Thamnovirus was expanded by two new species, Thamnovirus percae for Fiwi virus (FIWIV) and Thamnovi- rus kanderense for Kander virus (KNDV), both discovered by HTS in European perch (percid Perca fluviatilis Linnaeus, 1758) sampled in Switzerland [20] (TaxoProp 2021.011M.R .Filoviridae_1ngen_3nsp). Family Lispiviridae Family Lispiviridae was thoroughly revised (TaxoProp 2021.016M.R.Lispiviridae_16ngen_13nsp): • in genus Arlivirus, species Lishi arlivirus was renamed Arlivirus arachnae. Species Gerrid arlivirus, Hubei arlivirus, Odonate arlivirus, Tacheng arlivirus, and Wuchang arlivirus were renamed Sanstrivirus gerridis, Leocovirus coleopteris, Damravirus dentatis, Gani- avirus tachengense, and Nematovirus wuchangense, respectively, and moved into the newly created genera Sanstrivirus, Leocovirus, Damravirus, Ganiavirus, and Nematovirus, respectively; • genus Anicalvirus was created to include the new species Anicalvirus hangzhouense for Anisopteromalus caland- rae negative-strand RNA virus 2 (AcNSRV-2) discovered by HTS in a parasitoid wasp (pteromalid Anisoptero- malus calandrae (Howard, 1881)) sampled in Hángzhōu (杭州市), Zhèjiāng Province (浙江省), China [72]; • genus Anidravirus was created to include the new spe- cies Anidravirus hangzhouense for Anisopteromalus calandrae negative-strand RNA virus 1 (AcNSRV-1) discovered by HTS in parasitoid wasps (pteromalid Anisopteromalus calandrae (Howard, 1881)) sampled 2860 J. H. Kuhn et al. 1 3 humile (Mayr, 1868)) sampled in Catalonia (Cata- luña), Spain [68]; and o Synelinevirus bonnense for hymenopteran arli- related virus OKIAV98 (HARV-98) discovered by HTS in gall wasps (cynipid Synergus umbraculus Walker, 1874) sampled in Bonn, North Rhine-West- phalia (Nordrhein-Westfalen), Germany [27]; • genus Usmuvirus was created to include the new species Usmuvirus newyorkense for Amsterdam virus (AMSV) discovered by HTS in house mice (murid Mus musculus Linnaeus, 1758) sampled in New York, New York, USA [73]; and • genus Xenophyvirus was created to include the new spe- cies Xenophyvirus mathesonense for hemipteran arli- related virus OKIAV95 (HARV-95) discovered by HTS in moss bugs (peloridiid Xenophyes metoponcus Burck- hardt, 2011) sampled in Lake Matheson, South Island Westland District, New Zealand [27]. Family Mymonaviridae Genus Auricularimonavirus was expanded by one new species, Auricularimonavirus bondarzewiae, for Bondar- zewia berkeleyi negative-strand RNA virus 1 (BbNSRV-1) discovered by HTS in stump blossoms (bondarzewiaceaen Bondarzewia berkeleyi (Fr.) Bondartsev & Singer (1941)) [65] (TaxoProp 2021.041M.R.Mymonaviridae_5nsp). Genus Botrytimonavirus was expanded by one new spe- cies, Botrytimonavirus alphabotrytidis, for Botrytis cinerea negative-stranded RNA virus 5 (BcNSRV-5) and Sclerotinia sclerotiorum negative-stranded RNA virus 10 (SsNSRV-10) discovered in metatranscriptomes of sclerotiniaceaen fungi (Botrytis cinerea Pers. (1794) and Sclerotinia sclerotiorum (Lib.) de Bary (1884) sampled in Italy/Spain and China, respectively) [26, 53] (TaxoProp 2021.041M.R.Mymonaviri dae_5nsp). Sclerotinia sclerotiorum negative-stranded RNA virus 11 (SsNSRV-11) discovered in a Sclerotinia sclerotio- rum (Lib.) de Bary (1884) metatranscriptome dataset [26] was assigned to the already established species Botrytimo- navirus botrytidis (TaxoProp 2021.041M.R.Mymonavirid ae_5nsp). Genus Sclerotimonavirus was expanded by three new spe- cies (TaxoProp 2021.041M.R.Mymonaviridae_5nsp): • Sclerotimonavirus alphabotrytidis for Botrytis cinerea negative-stranded RNA virus 3 (BcNSRV-3) and Scle- rotinia sclerotiorum negative-stranded RNA virus 9 (SsNSRV-9) discovered in metatranscriptomes of scle- rotiniaceaen fungi (Botrytis cinerea Pers. (1794) and Sclerotinia sclerotiorum (Lib.) de Bary (1884) sampled in Italy/Spain and China, respectively) [26, 53]; • Sclerotimonavirus betabotrytidis for Botrytis cinerea negative-stranded RNA virus 4 (BcNSRV-4), discovered in the metatranscriptome of a sclerotiniaceaen fungus (Botrytis cinerea Pers. (1794)) sampled in Italy/Spain [53]; and • Sclerotimonavirus cryphonectriae for Cryphonectria parasitica sclerotimonavirus 1 (CpSV-1), discovered by HTS in a cryphonectriaceaen fungus (Cryphonectria parasitica (Murrill) M. E. Barr (1978)) sampled in Azer- baijan [18]. Family Nyamiviridae The names of all species in the family were changed to binomials as required by the recently amended ICVCN [25, 69, 81] (TaxoProp 2021.020M.R.Nyamiviridae_sprename). Genus Formivirus was expanded by one new species, Formivirus solenopsi, for Solenopsis invicta virus 15 (SoINV-15) identified in the transcriptome of red fire ants (myrmicine Solenopsis invicta Buren, 1972) sampled in the US and Taiwan [77] (TaxoProp 2021.019M.R.Nyamivirid ae_3nsp). Genus Nyavirus was expanded by two new species (Taxo- Prop 2021.019M.R.Nyamiviridae_3nsp): • Nyavirus argatis, for Sekira virus (SEKRV) identified in the RNA virome of a pool of ticks (argasid Argas japoni- cus Yamaguti, Clifford and Tipton, 1968) sampled in Mishima (三島市), Shizuoka Prefecture (静岡県), Japan [28]; and • Nyavirus somateriae for Jeremy Point nyavirus (JPNV) identified in the spleen of a common eider (anatid Soma- teria mollissima (Linnaeus, 1758)) collected at Jeremy Point, Massachusetts, USA, in 2013 (unpublished; Gen- Bank: MN045233). Family Paramyxoviridae Genus Jeilongvirus was expanded by eight new species (TaxoProp: 2021.014M.R.Jeilongvirus_8nsp): • Jeilongvirus anhuiense for Miniopterus schreibersii para- myxovirus (MisPV) discovered by HTS in Schreibers’s long-fingered bats (miniopterid Miniopterus schreibersii (Kuhl, 1817)) sampled in China [75]; • Jeilongvirus apodemi for rodent paramyxovirus (RoPV) discovered by HTS in a Korean field mouse (murid Apodemus peninsulae (Thomas, 1907)) sampled in China [76]; • Jeilongvirus comorosense for bat paramyxovirus 16797 (BatPV-1) identified in a Comoran long-fingered bat (miniopterid Miniopterus griveaudi Harrison, 1959) sam- pled in Comoros (unpublished; GenBank: MG203877); 2861Taxonomic update of phylum Negarnaviricota 1 3 • Jeilongvirus erinacei for belerina virus (BeV) discovered by HTS in West European hedgehogs (erinaceid Erina- ceus europaeus Linnaeus, 1758) sampled in Belgium [66]; • Jeilongvirus felis for feline paramyxovirus (FPaV) identi- fied in domestic cats (feline Felis catus Linnaeus, 1758) sampled in Germany [55, 61]; • Jeilongvirus madagascarense for bat paramyxovirus 17770 (BatPV-2) identified in a Comoran long-fin- gered bat (miniopterid Miniopterus griveaudi Harrison, 1959) sampled in Madagascar (unpublished; GenBank: MG203878); • Jeilongvirus murinae for bat paramyxovirus (BatPV- 3) identified in a greater tube-nosed bat (vespertilionid Murina leucogaster (Milne-Edwards, 1872) sampled in China [75]; and • Jeilongvirus rungweense for ruloma virus (RulV) detected by HTS in a Machangu’s brush-furred rat (murid Lophuromys machangui Verheyen, Hulselmans, Dierckx, Mulungu, Leirs, Corti & Verheyen, 2007) sampled in Rungwe District, Mbeya Region, Tanzania [67]. Family Rhabdoviridae The names of all species in the family were changed to binomials as required by the recently amended ICVCN [25, 69, 81] (TaxoProps 2021.036M.R.Rhabdoviridae_sprename, 2021.042M.R.Corrections_Riboviria_Duplodnaviria, and 2021.043M.R.Corrections_Mononegavirales_Bunyavirales). Five new genera were created and included in the subfam- ily Alpharhabdovirinae: • genus Amplylivirus, including one novel species, Amplylivirus cinereus, for frog lyssa-like virus 1 (FLLV- 1) detected in an American green tree frog (hylid Dryo- phytes cinereus (Schneider, 1799)) (unpublished; Gen- Bank: MK473367) (2021.035M.R.Rhabdoviridae_2nge n_2nsp); • genus Cetarhavirus for two new species (2021.003M.R. Alpharhabdovirinae_3ngen_7nsp): o Cetarhavirus laganorhynchus for dolphin rhabdovi- rus (DRV) isolated from a white-beaked dolphin (delphinid Lagenorhynchus albirostris (Gray, 1846)) sampled in Schiermonnikoog, Friesland, Holland [45]; and o Cetarhavirus phocoena for harbour porpoise rhab- dovirus (HPRV) isolated from a harbour porpoise (phocoenid Phocoena phocoena (Linnaeus, 1758)) stranded off the coast of Alaska [15]; • genus Replylivirus, including one novel species, Replyli- virus allogus, for anole lyssa-like virus 1 (ALLV-1) iden- tified in RNAseq data from a Spanish flag anole (dac- tyloid Anolis allogus Barbour & Ramsden, 1919) [22] (2021.035M.R.Rhabdoviridae_2ngen_2nsp); • genus Siniperhavirus for two new species (2021.003M.R .Alpharhabdovirinae_3ngen_7nsp): o Siniperhavirus chuatsi for Siniperca chuatsi rhab- dovirus (SRCV) isolated from mandarin fish (sin- ipercid Siniperca chuatsi (Basilewsky, 1855)) collected in Guǎngdōng Province (广东省 ), China [86, 87]; Chinese rice-field eel rhabdovirus (CrERV) isolated from Chinese ricefield eels (syn- branchid Monopterus albus (Zuiew, 1793)) sam- pled in Qiánjiāng (潜江市), Húběi Province (湖 北省), China [38]; hybrid snakehead rhabdovirus (HSHRV) isolated from hybrid snakehead (chan- nid Channa maculata × Channa argus) sampled in Fóshān (佛山市), Guǎngdōng Province (广东省), China [80]; and Micropterus salmoides rhabdovirus (MSRV) isolated from largemouth bass (centrarchid Micropterus salmoides (Lacépède, 1802)) sampled in Yúháng (余杭区), Zhèjiāng Province (浙江省), China [40]; and o Siniperhavirus zoarces for harbour eelpout rhab- dovirus (EPRV) detected by HTS in eelpout (zoarcid Zoarces viviparous (Linnaeus, 1758)) near Stock- holm, Sweden [2]; • genus Scophrhavirus for two new species (2021.003M.R.Alpharhabdovirinae_3ngen_7nsp): o Scophrhavirus chanodychthys for Wuhan redfin cul- ter dimarhabdovirus (WhRFCRV) detected by high- throughput sequencing in redfin culter (cyprinid Chanodichthys erythropterus (Basilewsky, 1855)) collected in “Húběi Province (湖北省)”, China [58]; and o Scophrhavirus maximus for Scophthalmus maxi- mus rhabdovirus (SMRV) isolated from turbot fish (scophthalmid Scophthalmus maximus (Linnaeus, 1758)) sampled in Shāndōng Province (山东省), China [82]. Genus Alphanucleorhabdovirus was expanded by one new species, Alphanucleorhabdovirus joa, for joa yellow blotch associated virus (JYBaV) identified in joá (solanace- aen Solanum aculeatissimum L.) plants sampled in Manaus, State of Amazonas (Estado de Amazonas), Brazil [11] (Tax- oProp 2021.001M.A.v1.Betarhabdovirinae_7nsp). Genus Betanucleorhabdovirus was expanded by one new species, Betanucleorhabdovirus bacopae, for Bacopa mon- nieri virus 2 (BmV-2) identified in transcriptomic data of water hyssop (plantaginaceaen Bacopa monnieri L.) tissues 2862 J. H. Kuhn et al. 1 3 sampled in Lucknow, Uttar Pradesh, India [60] (TaxoProp 2021.001M.A.v1.Betarhabdovirinae_7nsp). Genus Cytorhabdovirus was expanded by five new spe- cies (TaxoProp 2021.001M.A.v1.Betarhabdovirinae_7nsp): • Cytorhabdovirus bacopae for Bacopa monnieri virus 1 (BmV-1) identified in transcriptomic data of water hys- sop (plantaginaceaen Bacopa monnieri L.) tissues sam- pled in Lucknow, Uttar Pradesh, India [60]; • Cytorhabdovirus chrysanthemi for chrysanthemum yel- low dwarf associated virus (CYDaV) identified in grey- leaved euryops (asteraceaen Euryops pectinatus (L.) Cass.) plant sampled in Chóngqìng (重庆市), China [37]; • Cytorhabdovirus cucurbitae for cucurbit cytorhabdovi- rus 1 (CuCV-1) identified in zucchini (cucurbitaceaen Cucurbita pepo L.) plants sampled in Greece [44]; • Cytorhabdovirus broussonetiae for paper mulberry mosaic associated virus (PMuMaV) identified in paper mulberry (moraceaen Broussonetia papyrifera L.) plants sampled in Chóngqìng (重庆市), China [51]; and • Cytorhabdovirus rosae for rose virus R (RVR), identified in a rose ‘Hugh Dickson’ plant from Maryland, USA [5]. Genus Hapavirus was expanded by two new species (TaxoProp 2021.004M.R.Alpharhabdovirinae_13nsp): • Hapavirus bangoran for Bangoran virus (BGNV) found in mosquitoes (culicid Culex perfuscus Edwards, 1914) sampled in Bangoran, Bamingui-Bangoran Prefecture, Central African Republic [6, 14]; and • Hapavirus porton for Porton virus (PORV) found in mos- quitoes (culicid Mansonia uniformis (Theobald, 1901)) sampled in Sarawak state, Malaysia [6]. Genus Ledantevirus was expanded by one new species, Ledantevirus taiyi, for Taiyi bat virus (TYBV) found in Chi- nese rufous horseshoe bats (rhinolophid Rhinolophus sinicus K. Andersen, 1905) sampled in Xiánníng (咸宁市), Húběi Province (湖北省), China [39] (TaxoProp 2021.004M.R.A lpharhabdovirinae_13nsp). Genus Perhabdovirus was expanded by one new species, Perhabdovirus leman, for Leman virus (LeRV) isolated from perch fry sampled from Lake Geneva (Lac Léman), France [47] (TaxoProp 2021.003M.R.Alpharhabdovirinae_3ngen_7nsp). Genus Sunrhavirus was expanded by eight new species (TaxoProp 2021.004M.R.Alpharhabdovirinae_13nsp): • Sunrhavirus sandjimba for Sandjimba virus (SJAV) found in sedge warblers (acrocephalid Acrocephalus schoenobaenus (Linnaeus, 1758)) collected at the Land- jia river, Ombella-M'Poko Prefecture, Central African Republic [6, 14]; • Sunrhavirus nasoule for Nasoule virus (NASV) found in little greenbuls (pycnonotid Eurillas virens (Cassin, 1857)) sampled in Nasoule, Mambéré-Kadéi Prefecture, Central African Republic [6, 14]; • Sunrhavirus boteke for Boteke virus (BOTV) found in mosquitoes (culicid Coquillettidia maculipennis (Theobald, 1911)) sampled in Boteke, Ombella-M'Poko Prefecture, Central African Republic [6, 14]; • Sunrhavirus bimbo for Bimbo virus (BBOV) found in yellow-crowned bishop (ploceid Euplectes afer (Gmelin, 1789)) sampled in Kolongo, Ombella-M'Poko Prefecture, Central African Republic [6, 14]; • Sunrhavirus kolongo for Kolongo virus (KOLV) found in yellow-crowned bishop (ploceid Euplectes afer (Gmelin, 1789)) sampled in Bangui, Central African Republic [6, 14]; • Sunrhavirus ouango for Ouango virus (OUAV) found in black-headed weaves (ploceid Ploceus melanocephalus (Linnaeus, 1758)) collected at the Landjia river, Ombella- M'Poko Prefecture, Central African Republic [6, 14]; • Sunrhavirus alexandria for Burg el Arab virus (BEAV) found in lesser whitethroats (sylviid Curruca curruca (Linnaeus, 1758)) sampled in Borg El Arab/Burj al-'Arab, Alexandria Governorate, Egypt [6]; and • Sunrhavirus matariya for Matariya virus (MTYV) found in lesser whitethroats (sylviid Curruca curruca (Linnaeus, 1758)) sampled in Port Saïd/Būrsaʿīd, Port Saïd Governorate, Egypt [6]. Genus Vesiculovirus was expanded by two new species (TaxoProp 2021.004M.R.Alpharhabdovirinae_13nsp): • Vesiculovirus mediterranean for Mediterranean bat virus (MBV) found in bats sampled in Algeria, Morocco, and Spain (unpublished; GenBank: MW557331); and • Vesiculovirus yinshui for Yinshui bat virus (YSBV) found in Chinese rufous horseshoe bats (rhinolophid Rhinolo- phus sinicus K. Andersen, 1905) sampled in Xiánníng (咸宁市), Húběi Province (湖北省), China [39]. Family Sunviridae The name of the only species in the family was changed to a binomial as required by the recently amended ICVCN [25, 69, 81] (2021.037M.R.Sunviridae_sprename). Family Xinmoviridae Family Xinmoviridae was thoroughly revised (2021.039M.R.Xinmoviridae_11ngen_8nsp): • in the genus Anphevirus, the species Shuangao anphevi- rus was abolished, and the species Xincheng anphevirus was renamed Anphevirus xinchengense. Species Dipteran anphevirus, Odonate anphevirus, Drosophilid anphevi- 2863Taxonomic update of phylum Negarnaviricota 1 3 rus, Bolahun anphevirus, and Orthopteran anphevirus were renamed Alasvirus muscae, Draselvirus dentati, Drunivirus chambonense, Gambievirus bolahunense, and Hoptevirus orthopteris, respectively, and moved into the newly created genera Alasvirus, Draselvirus, Drunivirus, Gambievirus, and Hoptevirus, respectively; • genus Doupovirus was created to include the new species Doupovirus australiaense for Culex mononega-like virus 2 (CMLV-2) discovered by HTS in mosquitoes (culicid Culex australicus Dobrotworsky & Drummond, 1953) sampled in Point Douro, Western Australia, Australia [57]; • genus Gambievirus was created to include the new spe- cies Gambievirus senegalense for Gambie virus (GAMV) discovered by HTS in mosquitoes (culicid Anopheles gambiae Giles, 1902) sampled in Senegal [17]; • genus Gylbovirus was created to include new species Gylbovirus aagae for Aedes anphevirus (AeAV) discov- ered by HTS in yellow fever mosquitoes (culicid Aedes aegypti (Linnaeus in Hasselquist, 1762)) and the Aag2 Aedes albopictus cell line [10, 48]; • genus Madalivirus was created to include two new spe- cies: o Madalivirus amapaense for Anopheles marajoara virus (AnMV) discovered by HTS in mosquitoes (culicid Anopheles marajoara Galvao & Damesceno 1942) sampled in Macapá, Amapá State, Brazil [56]; and o Madalivirus amazonaense for Anopheles darlingi virus (AnDV) discovered by HTS in American malaria mosquitoes (culicid Anopheles darlingi Root, 1926) sampled in Macapá, Amapá State, and Manaus, Amazonas State, Brazil [56]; • genus Pelmivirus was created to include new species Pelmivirus eymattense for hymenopteran anphe-related virus OKIAV71 (HARV-71) discovered by HTS in para- sitoid wasps (ichneumonid Heteropelma amictum (Fab- ricius, 1775)) sampled in Eymatt, Bern, Switzerland [27]; • genus Triniovirus was created to include the new species Trin- iovirus yonagoense for Culex tritaeniorhynchus anphevirus (CtAV) discovered by HTS in mosquitoes (culicid Culex tri- taeniorhynchus Giles, 1901) sampled in Yonago (米子市), Tottori Prefecture (鳥取県), Japan [16]; and • genus Ulegvirus was created for new species Ulegvi- rus freckenfeldense for odonatan anphe-related virus OKIAV59 (OARV-59) discovered by HTS in golden- ringed dragonflies (cordulegastrid Cordulegaster boltonii (Donovan, 1807)) sampled in Freckenfeld, Rhineland- Palatinate (Rheinland-Pfalz), Germany [27]. Yunchangviricetes The names of all species in the class were changed to bino- mials as required by the recently amended International Code of Virus Classification and Nomenclature (ICVCN) [25, 69, 81] (TaxoProp 2021.018M.R.Negarnaviricota _sprename). Taxonomic changes in subphylum Polyploviricotina Taxonomic changes within order Articulavirales (Polyploviricotina: Insthoviricetes) Family Amnoonviridae The name of the only species in the family was changed to a binomial as required by the recently amended International Code of Virus Classification and Nomenclature (ICVCN) [25, 69, 81] (TaxoProp 2021.018M.R.Negarnaviricota_sprename). Family Orthomyxoviridae The names of all species in the family were changed to binomials as required by the recently a m e n d e d I C VC N [ 2 5 , 6 9 , 8 1 ] ( Ta xo P ro p 2021.024M.R.Orthomyxoviridae_sprename). One new genus, Mykissvirus, was established to include one new species, Mykissvirus tructae, for rain- bow trout orthomyxovirus (RbtOV) and steelhead trout orthomyxovirus (SttOV-1) first isolated from commer- cially reared rainbow trout (salmonid Oncorhynchus mykiss (Walbaum, 1792)) in Hagerman Valley, Idaho, USA [4] (TaxoProp 2021.022M.R.Orthomyxoviridae_1 ngen_1nsp_Mykiss). One new genus, Sardinovirus, was established to include one new species, Sardinovirus pilchardi, for pilchard orthomyxovirus (POMV) first isolated from blue pilchards (clupeid Sardinops sagax (Jenyns, 1842)) sampled off the coast of South Australia and subsequently Tasmania [19, 41] (TaxoProp 2021.023M.R.Orthomyxoviridae_1ngen_1 nsp_Sardino). Genus Quaranjavirus was expanded by four new species (TaxoProp 2021.034M.R.Quaranjavirus_4nsp): • Quaranjavirus araguariense for Araguari virus (ARAV) isolated from a gray four-eyed opossum (didelphid Phi- lander opossum (Linnaeus, 1758)) sampled in Serra do Navio, Amapá State, Brazil [9, 79]; • Quaranjavirus chadense for Lake Chad virus (LKCV) isolated from a vitelline masked weaver (ploceid Ploceus 2864 J. H. Kuhn et al. 1 3 vitellinus (Lichtenstein, 1823)), sampled at Lake Chad, Nigeria [50]; • Quaranjavirus tyulekense for Tyulek virus (TLKV) isolated from ticks (argasid Argas vulgaris Filippova, 1961) sampled near Tûlëk/Tölök (Tюлёк/Tөлөк), Naryn Region (Hapынcкaя oблacть), Kirghiz Soviet Socialist Republic, USSR [85]; and • Quaranjavirus wellfleetense for Wellfleet Bay virus (WFBV) isolated from common eiders (anatid Somate- ria mollissima (Linnaeus, 1758)) sampled at Cape Cod, Massachusetts, USA [1]. Genus Thogotovirus was expanded by six new species (TaxoProp 2021.038M.R.Thogotovirus_6nsp): • Thogotovirus bourbonense for Bourbon virus (BRBV) isolated from a human sampled in Bourbon County, Kan- sas, USA [30]; • Thogotovirus josense for Jos virus (JOSV) isolated from a zebu (bovid Bos indicus Linnaeus, 1758) and ticks (ixodid Amblyomma variegatum (Fabricius, 1794) and Rhipicephalus decoloratus Koch, 1844)) sampled in Jos, Plateau State, Nigeria [36]; • Thogotovirus ozense for Oz virus (OZV) isolated from ticks (ixodid Amblyomma testudinarium Koch, 1844) sampled in Ehime Prefecture (愛媛県), Japan [13]; • Thogotovirus sinuense for Sinu virus (SINUV) isolated from a pool of mosquitoes collected in San Bernardo del Viento, Córdoba Department, Colombia [8]; • Thogotovirus thailandense for Thailand tick thogotovirus (TT-THOV) detected by HTS in ticks (ixodid Boophilus sp.) sampled in Tha Wang Pha District (ท่าวังผา), Nan Province (น่าน), Thailand [63]; and • Thogotovirus upoluense for Upolu virus (UPOV) isolated from ticks (ixodid Ornithodoros capensis Neumann, 1901) sampled on Upolu Cay, Great Barrier Reef, Aus- tralia [12]. Taxonomic changes within order Bunyavirales (Polyploviricotina: Ellioviricetes) The order Bunyavirales was expanded by two new families (2021.040M.R.Bunyavirales_2nfam_2ngen_6nsp): • Discoviridae: genus Orthodiscovirus was created for five species: o Orthodiscovirus coniellae for Coniothyrium diplo- diella negative-stranded RNA virus 1 (CdNSRV-1) and Plasmopara viticola lesion associated mycobun- yavirales-like virus 9, discovered by HTS in an isolate of a fungus (melanconidaceaen Coniella diplodiella (Speg.) Petr. & Syd., (1927)) and in plant lesions caused by grapevine downy mildew (peronosporace- aen Plasmopara viticola (Berk. & M.A. Curtis) Berl. & De Toni, (1888)), respectively [42]; o Orthodiscovirus hispaniae for Plasmopara viticola lesion associated mycobunyavirales-like virus 8 (PvLAM-LV8) discovered by HTS in plant lesions caused by grapevine downy mildew (peronosporace- aen Plasmopara viticola (Berk. & M.A. Curtis) Berl. & De Toni, (1888)) [7]; o Orthodiscovirus iberiae for Plasmopara viticola lesion associated mycobunyavirales-like virus 4 (PvLAM-LV4) discovered by HTS in plant lesions caused by grapevine downy mildew (peronosporace- aen Plasmopara viticola (Berk. & M.A. Curtis) Berl. & De Toni, (1888)) [7]; o Orthodiscovirus missouriense for Penicillium disco- virus (PDV) discovered by HTS in a fungus (tricho- comaceaen Penicillium atramentosum Thom, C. 1910) [31]; and o Orthodiscovirus penicillii for Penicillium roseopur- pureum negative ssRNA virus 1 (PrNssV-1) discov- ered by HTS in a fungus (trichocomaceaen Penicil- lium roseopurpureum Dierckx, R.P. 1901) [42]. • Tulasviridae: genus Orthotulasvirus was created to include one species, Orthotulasvirus tulasnellae, for Tulasnella bunyavirales-like virus 1 (TB-LV) discovered by HTS in an orchid mycorrhizal virome and confirmed to be present in an isolate of an effused fungus (tulasnel- laceaen Tulasnella sp.) by full length Sanger sequencing of its cDNA [62]. Family Arenaviridae Genus Mammarenavirus was expanded by two new species: • Mammarenavirus bituense for Bitu virus (BITV) detected by HTS in Namaqua micaelamys (Micaelamys namaq- uensis (A. Smith, 1834)) sampled in Angola (TaxoProp 2021.006M.R.Mammarenavirus_1nsp_Bitu) [64]; and • Mammarenavirus kwanzaense, for Kwanza virus (KWAV) detected by HTS in gray-bellied mice (Mus triton (Thomas, 1909)) sampled in Angola (TaxoProp 2021.007M.R.Mammarenavirus_1nsp_Kwanza) [64]. Family Fimoviridae The names of all species in the family were changed to binomials as required by the recently amended ICVCN [25, 69, 81] (TaxoProp 2021.013P.R.Fimoviridae_rename). Genus Emaravirus was expanded by three new species: 2865Taxonomic update of phylum Negarnaviricota 1 3 • Emaravirus aceris for maple mottle-associated emara- virus (MaMaV) discovered by HTS in maple (sap- indaceaen Acer sp.) sampled in Germany (TaxoProp 2021.010P.R.Emaravirus_1ns) [54]; • Emaravirus chrysanthemi for chrysanthemum mosaic-associated emaravirus (ChMaV) discov- ered by HTS in chrysanthemum (asteraceaen Chry- santhemum morifolium L.) sampled in Japan (TaxoProps 2021.011P.R.Emaravirus_1ns and 2021.043M.R.Corrections_Mononegavirales_Bunyavi- rales) [32]; and • Emaravirus quercus for common oak ringspot-associated emaravirus (CORaV) discovered by HTS in oak (fagace- aen Quercus robur L.) sampled in Germany and Scandi- navia (TaxoProp 2021.012P.R.Emaravirus_1ns) [52]. Family Hantaviridae The species name Seewis orhtohantavirus was corrected to Seewis orthohantavirus (TaxoProp 2021.042M.R.Corrections_Riboviria_Duplodnaviria). The species name Thottopalayam thottimvirus was corrected to Thottapalayam thottimvirus (TaxoProp 2021.042M.R.Corrections_Riboviria_Duplodnaviria). Family Leishbuviridae The name of the only species in the fam- ily was changed to a binomial as required by the recently amended ICVCN [25, 69, 81] (TaxoProp 2021.018M.R.Negarnaviricota_sprename). Table 1 ICTV-accepted taxonomy of the order Muvirales (Negarnaviricota: Haploviricotina: Chunqiuviricetes) as of March 2022 Note that viruses are real objects that are assigned to concepts that are called taxa. Species, genera, fami- lies, and orders are taxa ¶ Taxon names are always italicized and always begin with a capital letter & Virus names are not italicized and are not capitalized, except if the name or a name component is a proper noun. This column lists the virus names with their correct (lack of) capitalization Genus Species¶ Virus (Abbreviation)& Family Qinviridae Yingvirus Yingvirus beihaiense Běihǎi sesarmid crab virus 4 (BhSCV-4) Yingvirus charybdis Wēnzhōu qínvirus-like virus 2 (WzQLV-2) Yingvirus hubeiense Húběi qínvirus-like virus 1 (HbQLV-1) Yingvirus sanxiaense Sānxiá qínvirus-like virus 1 (SxQLV-1) Yingvirus shaheense Shāhé qínvirus-like virus 1 (ShQLV-1) Yingvirus wenzhouense Wēnzhōu qínvirus-like virus 1 (WzQLV-1) Yingvirus wuhanense Wǔhàn insect virus 15 (WhIV-15) Yingvirus xinzhouense Xīnzhōu nematode virus 3 (XzNV-3) Table 2 ICTV-accepted taxonomy of the order Serpentovirales (Negarnaviricota: Haploviricotina: Milneviricetes) as of March 2022 Note that viruses are real objects that are assigned to concepts that are called taxa. Species, genera, fami- lies, and orders are taxa ¶ Taxon names are always italicized and always begin with a capital letter & Virus names are not italicized and are not capitalized, except if the name or a name component is a proper noun. This column lists the virus names with their correct (lack of) capitalization Genus Species¶ Virus (Abbreviation)& Family Aspiviridae Ophiovirus Ophiovirus citri citrus psorosis virus (CPsV) Ophiovirus freesiae freesia sneak virus (FreSV) Ophiovirus lactucae lettuce ring necrosis virus (LRNV) Ophiovirus mirafioriense Mirafiori lettuce big-vein virus (MLBVV) Ophiovirus ranunculi ranunculus white mottle virus (RWMV) Ophiovirus tulipae tulip mild mottle mosaic virus (TMMMV) Ophiovirus vaccinii blueberry mosaic associated virus (BlMaV) 2866 J. H. Kuhn et al. 1 3 Table 3 ICTV-accepted taxonomy of the order Jingchuvirales (Negarnaviricota: Haploviricotina: Monjiviricetes) as of March 2022 Genus Species¶ Virus (Abbreviation)& Family Aliusviridae Obscuruvirus Obscuruvirus quintum Atrato chu-like virus 5 (AClV-5) Ollusvirus Ollusvirus coleopteri Húběi coleoptera virus 3 (HbCV-3) Ollusvirus culvertonense Culverton virus (CvV) Ollusvirus hanchengense Hánchéng leafhopper mivirus (HLMV) Ollusvirus hymenopteri hymenopteran chu-related virus 123 (HCrV-123) Ollusvirus insectii hymenopteran chu-related virus 126 (HCrV-126) Ollusvirus scaldisense Scaldis River bee virus (SRBV) Ollusvirus shayangense Shāyáng fly virus 1 (SyFV-1) Ollusvirus taiyuanense Tàiyuán leafhopper virus (TYLeV) Family Chuviridae Boscovirus Boscovirus hippoboscidae Wǔhàn louse fly virus 7 (WhLFV-7) Boscovirus hypoboscidae Wǔhàn louse fly virus 6 (WhLFV-6) Chuvivirus Chuvivirus brunnichi Wēnlǐng crustacean virus 14 (WlCV-14) Chuvivirus canceris Wēnzhōu crab virus 2 (WzCV-2) Culicidavirus Culicidavirus culicidae Wǔhàn mosquito virus 8 (WhMV-8) Culicidavirus culicis Culex mosquito virus 5 (ClMV-5) Culicidavirus imjinense Imjin River virus 1 (IjRV-1) Culicidavirus quitotaense Culex mosquito virus 4 (ClMV-4) Demapteravirus Demapteravirus dermapteri dermapteran chu-related virus 142 (DCrV-142) Doliuvirus Doliuvirus culisetae Mos8Chu0 chuvirus (MoCV) Mivirus Mivirus amblyommae lone star tick chuvirus 1 (LSTCV-1) Mivirus boleense Bólè tick virus 3 (BTV-3) Mivirus changpingense Chāngpíng tick virus 2 (CpTV-2) Mivirus dermacentoris Chāngpíng tick virus 3 (CpTV-3) Tǎchéng tick virus 5 (TcTV-5) Mivirus genovaense Genoa virus (GeV) Mivirus karukeraense Karukera tick virus (KtV) Mivirus rhipicephali brown dog tick mivirus 1 (BDTMV-1) Mivirus suffolkense Suffolk virus (SFKV) Mivirus wuhanense Wǔhàn tick virus 2 (WhTV-2) Morsusvirus Morsusvirus argatis Tǎchéng tick virus 4 (TcTV-4) Nigecruvirus Nigecruvirus ixodes blacklegged tick chuvirus-2 (BlTC-2) Odonatavirus Odonatavirus draconis odonatan chu-related virus 137 (OCrV-137) Odonatavirus fabricii Húběi odonate virus 11 (HbOV-11) Odonatavirus odontis odonatan chu-related virus 136 (OCrV-136) Pediavirus Pediavirus cirripedis Běihǎi barnacle virus 9 (BhBV-9) Piscichuvirus Piscichuvirus franki Herr Frank virus 1 (HFrV-1) Piscichuvirus lycodontis Guǎngdōng red-banded snake chuvirus-like virus (GRSCV) Piscichuvirus sanxiaense Sānxiá atyid shrimp virus 4 (SxASC-4) Piscichuvirus wenlingense Wēnlǐng fish chu-like virus (WFClV) Pterovirus Pterovirus chulinense hymenopteran chu-related virus OKIAV147 (HCrV-147) Scarabeuvirus Scarabeuvirus blattae Wǔchāng cockroach virus 3 (WcLFV-3) Scarabeuvirus dentati Húběi chuvirus-like virus 3 (HbCLV-3) Scarabeuvirus hubeiense Húběi chuvirus-like virus 1 (HbCLV-1) Scarabeuvirus lampyris Lampyris noctiluca chuvirus-like virus 1 (LNClV-1) Scarabeuvirus lishiense Lĭshì spider virus 1 (LsSV-1) Taceavirus Taceavirus wenlingense Wēnlǐng crustacean virus 15 (WlCV-15) 2867Taxonomic update of phylum Negarnaviricota 1 3 Family Mypoviridae The name of the only species in the fam- ily was changed to a binomial as required by the recently amended ICVCN [25, 69, 81] (TaxoProp 2021.018M.R.Negarnaviricota_sprename). Family Peribunyaviridae Family Peribunyaviridae was expanded by three new genera (TaxoProp 2021.027M.R.Peribunyaviridae_3ngen_ 3nsp): • Khurdivirus, including one new species, Khurdivirus volgaense, for Khurdun virus (KHURV) isolated from a Eurasian coot (rallid Fulica atra Linnaeus, 1758) sam- pled in Astrakhan Region (Acтpaxaнcкaя oблacть), Rus- sia [83, 84]; • Lakivirus, including one new species, Lakivirus lakam- haense, for Lakamha virus (LAKV) discovered by HTS in a mosquito (culicid Wyeomyia complosa (Dyar, 1928)) sampled in Palenque, Chiapas, Mexico [29]; and • Lambavirus, including one new species, Lambavirus wis- consinense, for largemouth bass bunyavirus (LBBV) dis- covered by HTS in largemouth bass (centrarchid Microp- terus salmoides (Lacépède, 1802)) sampled in the upper Mississippi River, Wisconsin, USA [71]. The species name Lumbo orthbunyavirus was corrected to Lumbo orthobunyavirus (TaxoProp 2021.042M.R.Corrections_Riboviria_Duplodnaviria). Family Phasmaviridae The family was expanded by one new genus, Cicadellivi- rus, to include one new species, Cicadellivirus scaphoidei, for Scaphoideus titanus bunya-like virus 1 (StHV) (Taxo- Prop 2021.030M.R.Phasmaviridae_1ngen_5nsp) [46]. Genus Orthophasmavirus was expanded by four new species (TaxoProps 2021.030M.R.Phasmaviridae_1ngen_5nsp and 2021.043M.R.Corrections_Mononegavirales_Bunyavirales): • Orthophasmavirus coredoense for Coredo virus (CORV) discovered by HTS in mosquitoes (culicid Mansonia titil- lans (Walker, 1848)) sampled in Colombia (unpublished; GenBank: MN661021–3); • Orthophasmavirus barstukasense for Barstukas virus (BARV) discovered by HTS in mosquitoes (culicid Aedes sp.) sampled in California, USA [3]; • Orthophasmavirus miglotasense for Miglotas virus (MIGV) discovered by HTS in mosquitoes (culicid Aedes sp. and Culex sp.) sampled in California, USA [3]; and • Orthophasmavirus flenense for Flen virus (FLNV) dis- covered by HTS in mosquitoes (culicid Aedes cantans (Meigen, 1818)) sampled in Flen, Södermanland County (Södermanlands län), Sweden [43]. Family Wupedeviridae The name of the only species in the fam- ily was changed to a binomial as required by the recently amended ICVCN [25, 69, 81] (TaxoProp 2021.018M.R.Negarnaviricota_sprename). Summary A summary of the current, ICTV-accepted taxonomy of the phylum Negarnaviricota is presented in Tables 1, 2, 3, 4, 5, 6 and 7 and in the form of two posters (Supplementary Figure 1). Table 3 (continued) Genus Species¶ Virus (Abbreviation)& Family Crepuscuviridae Aqualaruvirus Aqualaruvirus sialis megalopteran chu-related virus 119 (MCrV-119) Family Myriaviridae Myriavirus Myriavirus myriapedis Húběi myriapoda virus 8 (HbMV-8) Family Natareviridae Charybdivirus Charybdivirus charybdis Wēnzhōu crab virus 3 (WzCV-3) Note that viruses are real objects that are assigned to concepts that are called taxa. Species, genera, families, and orders are taxa. ¶ Taxon names are always italicized and always begin with a capital letter. & Virus names are not italicized and are not capitalized, except if the name or a name component is a proper noun. This column lists the virus names with their correct (lack of) capitalization. Lists of viruses within a given species are provisional at this point and will likely be amended in the near future. 2868 J. H. Kuhn et al. 1 3 Table 4 ICTV-accepted taxonomy of the order Mononegavirales (Negarnaviricota: Haploviricotina: Monjiviricetes) as of March 2022 Genus Species¶ Virus (Abbreviation)& Family Artoviridae Hexartovirus Hexartovirus cirripedis Běihǎi barnacle virus 8 (BhBV-8) Hexartovirus lepeophtheiri Lepeophtheirus salmonis negative-stranded RNA virus 1 (LsNSRV-1) Peropuvirus Peropuvirus beihaiense Běihǎi rhabdo-like virus 1 (BhRLV-1) Peropuvirus dentati Húběi rhabdo-like virus 8 (HbRLV-8) Peropuvirus hubeiense Húběi rhabdo-like virus 6 (HbRLV-6) Peropuvirus juli Húběi rhabdo-like virus 5 (HbRLV-5) Peropuvirus lignarii Běihǎi rhabdo-like virus 2 (BhRLV-2) Peropuvirus melongenae Solanum melongena rhabdo-like virus (SmRLV) Peropuvirus pteromali Pteromalus puparum negative-strand RNA virus 1 (PpNSRV-1) Family Bornaviridae Carbovirus Carbovirus queenslandense jungle carpet python virus (JCPV) Carbovirus tapeti southwest carpet python virus (SWCPV) Cultervirus Cultervirus hemicultri Wǔhàn sharpbelly bornavirus (WhSBV) Orthobornavirus Orthobornavirus alphapsittaciforme parrot bornavirus 1 (PaBV-1) parrot bornavirus 2 (PaBV-2) parrot bornavirus 3 (PaBV-3) parrot bornavirus 4 (PaBV-4) parrot bornavirus 7 (PaBV-7) Orthobornavirus avisaquaticae aquatic bird bornavirus 1 (ABBV-1) aquatic bird bornavirus 2 (ABBV-2) Orthobornavirus betapsittaciforme parrot bornavirus 5 (PaBV-5) Orthobornavirus bornaense Borna disease virus 1 (BoDV-1) Borna disease virus 2 (BoDV-2) Orthobornavirus caenophidiae Caribbean watersnake bornavirus (CWBV) Mexican black-tailed rattlesnake bornavirus (MRBV) Orthobornavirus elapsoideae Loveridge’s garter snake virus 1 (LGSV-1) Orthobornavirus estrildidae estrildid finch bornavirus 1 (EsBV-1) Orthobornavirus sciuri variegated squirrel bornavirus 1 (VSBV-1) Orthobornavirus serini canary bornavirus 1 (CnBV-1) canary bornavirus 2 (CnBV-2) canary bornavirus 3 (CnBV-3) Family Filoviridae Cuevavirus Lloviu cuevavirus Lloviu virus (LLOV) Dianlovirus Mengla dianlovirus Měnglà virus (MLAV) Ebolavirus Bombali ebolavirus Bombali virus (BOMV) Bundibugyo ebolavirus Bundibugyo virus (BDBV) Reston ebolavirus Reston virus (RESTV) Sudan ebolavirus Sudan virus (SUDV) Tai Forest ebolavirus Taï Forest virus (TAFV) Zaire ebolavirus Ebola virus (EBOV) Marburgvirus Marburg marburgvirus Marburg virus (MARV) Ravn virus (RAVV) Oblavirus Oblavirus percae Oberland virus (OBLV) Striavirus Xilang striavirus Xīlǎng virus (XILV) Tapjovirus Tapjovirus bothropis Tapajós virus (TAPV) Thamnovirus Huangjiao thamnovirus Huángjiāo virus (HUJV) Thamnovirus kanderense Kander virus (KNDV) Thamnovirus percae Fiwi virus (FIWIV) 2869Taxonomic update of phylum Negarnaviricota 1 3 Table 4 (continued) Genus Species¶ Virus (Abbreviation)& Family Lispiviridae Anicalvirus Anicalvirus hangzhouense Anisopteromalus calandrae negative-strand RNA virus 2 (AcNSRV-2) Anidravirus Anidravirus hangzhouense Anisopteromalus calandrae negative-strand RNA virus 1 (AcNSRV-1) Arlivirus Arlivirus arachnae Lĭshì spider virus 2 (LsSV-2) Copasivirus Copasivirus ivindoense isopteran arli-related virus OKIAV103 (IARV-103) Copasivirus manlyvaleense Jimsystermes virus (JIMV) Cybitervirus Cybitervirus niederense coleopteran arli-related virus OKIAV107 (CARV-107) Damravirus Damravirus dentatis Húběi odonate virus 10 (HbOV-10) Ganiavirus Ganiavirus tachengense Tǎchéng tick virus 6 (TcTV-6) Leocovirus Leocovirus coleopteris Húběi rhabdo-like virus 3 (HbRLV-3) Nematovirus Nematovirus wuchangense Wǔchāng romanomermis nematode virus 2 (WcRNV-2) Phelinovirus Phelinovirus aphidis hymenopteran arli-related virus OKIAV99 (HARV-99) Rivapovirus Rivapovirus aleyrodidae hemipteran arli-related virus OKIAV94 (HARV-94) Sanstrivirus Sanstrivirus gerridis Sānxiá water strider virus 4 (SxWSV-4) Stylovirus Stylovirus niederense strepsipteran arli-related virus OKIAV104 (SARV-104) Supelovirus Supelovirus thailandense blattodean arli-related virus OKIAV102 (BARV-102) Synelinevirus Synelinevirus bonnense hymenopteran arli-related virus OKIAV98 (HARV-98) Synelinevirus paranaense Linepithema humile rhabdo-like virus 1 (LhuRLV-1) Usmuvirus Usmuvirus newyorkense Amsterdam virus (AMSV) Xenophyvirus Xenophyvirus mathesonense hemipteran arli-related virus OKIAV95 (HARV-95) Family Mymonaviridae Auricularimonavirus Auricularimonavirus auriculariae Auricularia heimuer negative-stranded RNA virus 1 (AhNRSV-1) Auricularimonavirus bondarzewiae Bondarzewia berkeleyi negative-strand RNA virus 1 (BbNSRV-1) Botrytimonavirus Botrytimonavirus alphabotrytidis Botrytis cinerea negative-stranded RNA virus 5 (BcNSRV-5) Sclerotinia sclerotiorum negative-stranded RNA virus 10 (SsNSRV-10) Botrytimonavirus botrytidis Botrytis cinerea negative-stranded RNA virus 7 (BcNSRV-7) Sclerotinia sclerotiorum negative-stranded RNA virus 11 (SsNSRV-11) Botrytimonavirus glycinis soybean leaf-associated negative-stranded RNA virus 3 (SLaNSRV-3) Botrytimonavirus sclerotiniae Sclerotinia sclerotiorum negative-stranded RNA virus 2 (SsNSRV-2) Sclerotinia sclerotiorum negative-stranded RNA virus 2-A (SsNSRV-2-A) Sclerotinia sclerotiorum negative-stranded RNA virus 4 (SsNSRV-4) Sclerotinia sclerotiorum negative-stranded RNA virus 4-A (SsNSRV-4-A) Hubramonavirus Hubramonavirus hubeiense Húběi rhabdo-like virus 4 (HbRLV-4) Hubramonavirus terrae H2BulkLitter1223 virus Lentimonavirus Lentimonavirus lentinulae Lentinula edodes negative-strand RNA virus 1 (LeNSRV-1) Penicillimonavirus Penicillimonavirus alphapenicillii Penicillium adametzioides negative-stranded RNA virus 1 (PaNsRV-1) Penicillimonavirus alphaplasmoparae Plasmopara viticola lesion associated mononegaambi virus 1 (PvaMV-1) Penicillimonavirus betapenicillii Penicillium glabrum negative-stranded RNA virus 1 (PgRlV-1) Penicillimonavirus betaplasmoparae Plasmopara viticola lesion associated mononegaambi virus 2 (PvaMV-2) Plasmopara viticola lesion associated mononegaambi virus 4 (PvaMV-4) Penicillimonavirus deltaplasmoparae Plasmopara viticola lesion associated mononegaambi virus 5 (PvaMV-5) Penicillimonavirus epsilonplasmoparae Plasmopara viticola lesion associated mononegaambi virus 6 (PvaMV-6) Penicillimonavirus etaplasmoparae Plasmopara viticola lesion associated mononegaambi virus 9 (PvaMV-9) Penicillimonavirus gammaplasmopara Plasmopara viticola lesion associated mononegaambi virus 3 (PvaMV-3) Penicillimonavirus kilnbarnense Kiln Barn virus (KBV) Penicillimonavirus zetaplasmoparae Plasmopara viticola lesion associated mononegaambi virus 7 (PvaMV-7) Plasmopamonavirus Plasmopamonavirus plasmoparae Plasmopara viticola lesion associated mononegaambi virus 8 (PvaMV-8) Phyllomonavirus Phyllomonavirus gysingense Gysinge virus (GYSV) Phyllomonavirus phyllospherae soybean leaf-associated negative-stranded RNA virus 4 (SLaNSRV-4) 2870 J. H. Kuhn et al. 1 3 Table 4 (continued) Genus Species¶ Virus (Abbreviation)& Rhizomonavirus Rhizomonavirus mali apple virus B (APPv-B) Sclerotimonavirus Sclerotimonavirus alphabotrytidis Botrytis cinerea negative-stranded RNA virus 3 (BcNSRV-3) Sclerotinia sclerotiorum negative-stranded RNA virus 9 (SsNSRV-9) Sclerotimonavirus alphaclarireediae Sclerotinia homoeocarpa TSA contig 1 (ShTSA-1) Sclerotinia homoeocarpa TSA contig 2 (ShTSA-2) Sclerotimonavirus alphaplasmoparae Plasmopara viticola lesion associated mymonavirus 1 (PvaMV-1) Sclerotimonavirus alternariae Alternaria tenuissima negative-stranded RNA virus 1 (AtNsRV-1) Sclerotimonavirus betabotrytidis Botrytis cinerea negative-stranded RNA virus 4 (BcNSRV-4) Sclerotimonavirus betaclarireediae Sclerotinia homoeocarpa TSA contig 3 (ShTSA-3) Sclerotimonavirus betaplasmoparae Plasmopara viticola lesion associated mononega virus 2 (PvaMV-2) Sclerotimonavirus botrytidis Botrytis cinerea mymonavirus 1 (BcMyV-1) Sclerotinia sclerotiorum negative-stranded RNA virus 7 (SsNSRV-7) Sclerotimonavirus cryphonectriae Cryphonectria parasitica sclerotimonavirus 1 (CpSV-1) Sclerotimonavirus fusarii Fusarium graminearum negative-stranded RNA virus 1 (FgNSRV-1) soybean leaf-associated negative-stranded RNA virus 1 (SLaNSRV-1) Sclerotimonavirus illinoisense soybean leaf-associated negative-stranded RNA virus 2 (SLaNSRV-2) Sclerotimonavirus penicillii Penicillium cairnsense negative-stranded RNA virus 1 (PcNsRV-1) Sclerotimonavirus sclerotiniae Sclerotinia sclerotiorum negative-stranded RNA virus 1 (SsNSRV-1) Sclerotinia sclerotiorum negative-stranded RNA virus 1-A (SsNSRV-1-A) Sclerotinia sclerotiorum negative-stranded RNA virus 3 (SsNSRV-3) Sclerotinia sclerotiorum negative-stranded RNA virus 3-A (SsNSRV-3-A) Sclerotimonavirus terrae H4BulkLitter234 virus Family Nyamiviridae Berhavirus Berhavirus beihaiense Běihǎi rhabdo-like virus 4 (BhRLV-4) Berhavirus radialis Běihǎi rhabdo-like virus 5 (BhRLV-5) Berhavirus sipunculi Běihǎi rhabdo-like virus 3 (BhRLV-3) Crustavirus Crustavirus beihaiense Běihǎi rhabdo-like virus 6 (BhRLV-6) Crustavirus wenlingense Wēnlǐng crustacean virus 12 (WlCV-12) Crustavirus wenzhouense Wēnzhōu crab virus 1 (WzCV-1) Formivirus Formivirus angliae Formica fusca virus 1 (FfusV-1) Formivirus chalybii hymenopteran orino-related virus OKIAV87 (HORV-87) Formivirus finnoniae Formica exsecta virus 4 (FeV-4) Formivirus gorytis hymenopteran orino-related virus OKIAV85 (HORV-85) Formivirus solenopsi Solenopsis invicta virus 15 (SoINV-15) Nyavirus Nyavirus argatis Sekira virus (SEKRV) Nyavirus midwayense Midway virus (MIDWV) Nyavirus nyamaniniense Nyamanini virus (NYMV) Nyavirus sanjacintoense San Jacinto virus (SJCV) Nyavirus sierranevadaense Sierra Nevada virus (SNVV) Nyavirus somateriae Jeremy Point nyavirus (JPNV) Orinovirus Orinovirus pasiphilae Orinoco virus (ONCV) Socyvirus Socyvirus heteroderae soybean cyst nematode virus 1 (SbCNV-1) Tapwovirus Tapwovirus cesti Wēnzhōu tapeworm virus 1 (WzTWV-1) Family Paramyxoviridae Subfamily Avulavirinae Metaavulavirus Avian metaavulavirus 2 avian paramyxovirus 2 (APMV-2) Avian metaavulavirus 5 avian paramyxovirus 5 (APMV-5) Avian metaavulavirus 6 avian paramyxovirus 6 (APMV-6) Avian metaavulavirus 7 avian paramyxovirus 7 (APMV-7) 2871Taxonomic update of phylum Negarnaviricota 1 3 Table 4 (continued) Genus Species¶ Virus (Abbreviation)& Avian metaavulavirus 8 avian paramyxovirus 8 (APMV-8) Avian metaavulavirus 10 avian paramyxovirus 10 (APMV-10) Avian metaavulavirus 11 avian paramyxovirus 11 (APMV-11) Avian metaavulavirus 14 avian paramyxovirus 14 (APMV-14) Avian metaavulavirus 15 avian paramyxovirus 15 (APMV-15) Avian metaavulavirus 20 avian paramyxovirus 20 (APMV-20) Avian metaavulavirus 22 avian paramyxovirus 22 (APMV-22) Orthoavulavirus Avian orthoavulavirus 1 avian paramyxovirus 1 (APMV-1)1 Avian orthoavulavirus 9 avian paramyxovirus 9 (APMV-9) Avian orthoavulavirus 12 avian paramyxovirus 12 (APMV-12) Avian orthoavulavirus 13 avian paramyxovirus 13 (APMV-13) Avian orthoavulavirus 16 avian paramyxovirus 16 (APMV-16) Avian orthoavulavirus 17 Antarctic penguin virus A (APV-A) Avian orthoavulavirus 18 Antarctic penguin virus B (APV-B) Avian orthoavulavirus 19 Antarctic penguin virus C (APV-C) Avian orthoavulavirus 21 avian paramyxovirus 21 (APMV-21) Paraavulavirus Avian paraavulavirus 3 avian paramyxovirus 3 (APMV-3) Avian paraavulavirus 4 avian paramyxovirus 4 (APMV-4) Subfamily Metaparamyxovirinae Synodonvirus Synodus synodonvirus Wēnlǐng triplecross lizardfish paramyxovirus (WTLPV) Subfamily Orthoparamyxovirinae Aquaparamyxovirus Oncorhynchus aquaparamyxovirus Pacific salmon paramyxovirus (PSPV) Salmo aquaparamyxovirus Atlantic salmon paramyxovirus (AsaPV) Ferlavirus Reptilian ferlavirus fer-de-lance virus (FDLV) Henipavirus Cedar henipavirus Cedar virus (CedV) Ghanaian bat henipavirus Ghana virus (GhV) Hendra henipavirus Hendra virus (HeV) Mojiang henipavirus Mòjiāng virus (MojV) Nipah henipavirus Nipah virus (NiV) Jeilongvirus Beilong jeilongvirus Beilong virus (BeiV) Jeilongvirus anhuiense Miniopterus schreibersii paramyxovirus (MisPV) Jeilongvirus apodemi rodent paramyxovirus (RoPV) Jeilongvirus comorosense bat paramyxovirus 16797 (BatPV-1) Jeilongvirus erinacei belerina virus (BeV) Jeilongvirus felis feline paramyxovirus (FPaV) Jeilongvirus madagascarense bat paramyxovirus 17770 (BatPV-2) Jeilongvirus murinae bat paramyxovirus (BatPV-3) Jeilongvirus rungweense ruloma virus (RulV) Jun jeilongvirus J virus (JV) Lophuromys jeilongvirus 1 Mount Mabu Lophuromys virus 1 (MMLV-1) Lophuromys jeilongvirus 2 Mount Mabu Lophuromys virus 2 (MMLV-2) Miniopteran jeilongvirus Shaan virus (ShaV) Myodes jeilongvirus Pohorje Myodes paramyxovirus 1 (PMPV-1) Tailam jeilongvirus Tailam virus (TaiV) Morbillivirus Canine morbillivirus canine distemper virus (CDV) Cetacean morbillivirus cetacean morbillivirus (CeMV) Feline morbillivirus feline morbillivirus (FeMV) Measles morbillivirus measles virus (MeV) Phocine morbillivirus phocine distemper virus (PDV) 2872 J. H. Kuhn et al. 1 3 Table 4 (continued) Genus Species¶ Virus (Abbreviation)& Rinderpest morbillivirus rinderpest virus (RPV) Small ruminant morbillivirus peste-des-petits-ruminants virus (PPRV) Narmovirus Mossman narmovirus Mossman virus (MossV) Myodes narmovirus bank vole virus 1 (BaV-1) Nariva narmovirus Nariva virus (NarV) Tupaia narmovirus Tupaia paramyxovirus (TupV) Respirovirus Bovine respirovirus 3 bovine parainfluenza virus 3 (BPIV-3) Caprine respirovirus 3 caprine parainfluenzavirus 3 (CPIV-3) Human respirovirus 1 human parainfluenza virus 1 (HPIV-1) Human respirovirus 3 human parainfluenza virus 3 (HPIV-3) Murine respirovirus Sendai virus (SeV) Porcine respirovirus 1 porcine parainfluenza virus 1 (PPIV-1) Squirrel respirovirus giant squirrel virus (GSqV) Salemvirus Salem salemvirus Salem virus (SalV) Subfamily Rubulavirinae Orthorubulavirus Human orthorubulavirus 2 human parainfluenza virus 2 (HPIV-2) Human orthorubulavirus 4 human parainfluenza virus 4a (HPIV-4a) human parainfluenza virus 4b (HPIV-4b) Mammalian orthorubulavirus 5 parainfluenza virus 5 (PIV-5) Mammalian orthorubulavirus 6 Alston virus (AlsV) Mapuera orthorubulavirus Mapuera virus (MapV) Mumps orthorubulavirus mumps virus (MuV) Porcine orthorubulavirus La Piedad Michoacán Mexico virus (LPMV) Simian orthorubulavirus simian virus 41 (SV-41) Pararubulavirus Achimota pararubulavirus 1 Achimota virus 1 (AchPV-1) Achimota pararubulavirus 2 Achimota virus 2 (AchPV-2) Hervey pararubulavirus Hervey virus (HerV) Menangle pararubulavirus Menangle virus (MenPV) Sosuga pararubulavirus Sosuga virus (SOSV) Teviot pararubulavirus Teviot virus (TevPV) Tioman pararubulavirus Tioman virus (TioPV) Tuhoko pararubulavirus 1 Tuhoko virus 1 (ThkPV-1) Tuhoko pararubulavirus 2 Tuhoko virus 2 (ThkPV-2) Tuhoko pararubulavirus 3 Tuhoko virus 3 (ThkPV-3) Unassigned (to subfamilies) Cynoglossusvirus Cynoglossus cynoglossusvirus Wēnlǐng tonguesole paramyxovirus (WTSPV) Hoplichthysvirus Hoplichthys hoplichthysvirus Wēnlǐng hoplichthys paramyxovirus (WHPV) Scoliodonvirus Scoliodon scoliodonvirus Wēnzhōu pacific spadenose shark paramyxovirus (WPSSPV) Family Pneumoviridae Metapneumovirus Avian metapneumovirus avian metapneumovirus (AMPV) Human metapneumovirus human metapneumovirus (HMPV) Orthopneumovirus Bovine orthopneumovirus bovine respiratory syncytial virus (BRSV) Human orthopneumovirus human respiratory syncytial virus (HRSV) Murine orthopneumovirus murine pneumonia virus (MPV) Family Rhabdoviridae Subfamily Alpharhabdovirinae Almendravirus Almendravirus almendras Puerto Almendras virus (PTAMV) Almendravirus arboretum Arboretum virus (ABTV) 2873Taxonomic update of phylum Negarnaviricota 1 3 Table 4 (continued) Genus Species¶ Virus (Abbreviation)& Almendravirus balsa Balsa virus (BALV) Almendravirus chico Rio Chico virus (RCHV) Almendravirus cootbay Coot Bay virus (CBV) Almendravirus menghai Menghai virus (MRV) Alphanemrhavirus Alphanemrhavirus xingshan Xingshan nematode virus 4 (XsNV-4) Alphanemrhavirus xinzhou Xinzhou nematode virus 4 (XzNV-4) Alphapaprhavirus Alphapaprhavirus hubei Hubei lepidoptera virus 2 (HbLV-2) Alphapaprhavirus pararge Pararge aegeria rhabdovirus (PAeRV) Alpharicinrhavirus Alpharicinrhavirus blanchseco Blanchseco virus (BCOV) Alpharicinrhavirus bole Bole tick virus 2 (BlTV-2) Alpharicinrhavirus wuhan Wuhan tick virus 1 (WhTV-1) Amplylivirus Amplylivirus cinereus frog lyssa-like virus 1 (FLLV-1) Arurhavirus Arurhavirus aruac Aruac virus (ARUV) Arurhavirus inhangapi Inhangapi virus (INHV) Arurhavirus santabarbara Santa Barbara virus (SBAV) Arurhavirus xiburema Xiburema virus (XIBV) Barhavirus Barhavirus bahia Bahia Grande virus (BGV) Harlingen virus (HARV) Barhavirus muir Muir Springs virus (MSV) Caligrhavirus Caligrhavirus caligus Caligus rogercresseyi rhabdovirus (CRogRV) Caligrhavirus lepeophtheirus Lepeophtheirus salmonis rhabdovirus 127 (LSalRV-127) Caligrhavirus salmonlouse Lepeophtheirus salmonis rhabdovirus 9 (LSalRV-9) Cetarhavirus Cetarhavirus laganorhynchus dolphin rhabdovirus (DRV) Cetarhavirus phocoena harbour porpoise rhabdovirus (HPRV) Curiovirus Curiovirus curionopolis Curionopolis virus (CURV) Curiovirus iriri Iriri virus (IRIRV) Curiovirus itacaiunas Itacaiunas virus (ITAV) Curiovirus rochambeau Rochambeau virus (RBUV) Ephemerovirus Ephemerovirus adelaide Adelaide River virus (ARV) Ephemerovirus berrimah Berrimah virus (BRMV) Ephemerovirus febris bovine ephemeral fever virus (BEFV) Ephemerovirus hayes Hayes Yard virus (HYV) Ephemerovirus kent New Kent County virus (NKCV) Ephemerovirus kimberley Kimberley virus (KIMV) Malakal virus (MALV) Ephemerovirus koolpinyah Koolpinyah virus (KOOLV) Ephemerovirus kotonkan kotonkan virus (KOTV) Ephemerovirus obodhiang Obodhiang virus (OBOV) Ephemerovirus puchong Puchong virus (PUCV) Ephemerovirus yata Yata virus (YATV) Hapavirus Hapavirus bangoran Bangoran virus (BGNV) Hapavirus flanders Flanders virus (FLAV) Hapavirus graylodge Gray Lodge virus (GLOV) Hapavirus hartpark Hart Park virus (HPV) Hapavirus holmes Holmes Jungle virus (HOJV) Hapavirus joinjakaka Joinjakaka virus (JOIV) Hapavirus kamese Kamese virus (KAMV) Hapavirus lajoya La Joya virus (LJV) Hapavirus landjia Landjia virus (LANV = LJAV) Hapavirus manitoba Manitoba virus (MANV = MNTBV) 2874 J. H. Kuhn et al. 1 3 Table 4 (continued) Genus Species¶ Virus (Abbreviation)& Hapavirus marco Marco virus (MCOV) Hapavirus mosqueiro Mosqueiro virus (MQOV) Hapavirus mossuril Mossuril virus (MOSV) Hapavirus ngaingan Ngaingan virus (NGAV) Hapavirus ord Ord River virus (ORV) Hapavirus parry Parry Creek virus (PCV) Hapavirus porton Porton virus (PORV) Hapavirus wongabel Wongabel virus (WONV) Ledantevirus Ledantevirus barur Barur virus (BARV) Ledantevirus bughendera Bughendera virus (BUGV) Ledantevirus elgon Mount Elgon bat virus (MEBV) Ledantevirus fikirini Fikirini virus (FKRV) Ledantevirus fukuoka Fukuoka virus (FUKV) Ledantevirus kanyawara Kanyawara virus (KYAV) Ledantevirus kern Kern Canyon virus (KCV) Ledantevirus keuraliba Keuraliba virus (KEUV) Ledantevirus kolente Kolente virus (KOLEV) Ledantevirus kumasi Kumasi rhabdovirus (KRV) Ledantevirus ledantec Le Dantec virus (LDV) Ledantevirus nishimuro Nishimuro virus (NISV) Ledantevirus nkolbisson Nkolbisson virus (NKOV) Ledantevirus oita Oita virus (OITAV) Ledantevirus taiyi Taiyi bat virus (TYBV) Ledantevirus vaprio Vaprio virus (VAPV) Ledantevirus wuhan Wuhan louse fly virus 5 (WLFV-5) Ledantevirus yongjia Yongjia tick virus 2 (YTV-2) Lostrhavirus Lostrhavirus hyalomma Xinjiang tick rhabdovirus (XjTRV) Lostrhavirus lonestar lone star tick rhabdovirus (LITRV) Lyssavirus Lyssavirus aravan Aravan virus (ARAV) Lyssavirus australis Australian bat lyssavirus (ABLV) Lyssavirus bokeloh Bokeloh bat lyssavirus (BBLV) Lyssavirus caucasicus West Caucasian bat virus (WCBV) Lyssavirus duvenhage Duvenhage virus (DUVV) Lyssavirus formosa Taiwan bat lyssavirus (TWBLV) Lyssavirus gannoruwa Gannoruwa bat lyssavirus (GBLV) Lyssavirus hamburg European bat lyssavirus 1 (EBLV-1) Lyssavirus helsinki European bat lyssavirus 2 (EBLV-2) Lyssavirus ikoma Ikoma lyssavirus (IKOV) Lyssavirus irkut Irkut virus (IRKV) Lyssavirus khujand Khujand virus (KHUV) Lyssavirus lagos Lagos bat virus (LBV) Lyssavirus lleida Lleida bat lyssavirus (LLEBV) Lyssavirus mokola Mokola virus (MOKV) Lyssavirus rabies rabies virus (RABV) Lyssavirus shimoni Shimoni bat virus (SHIBV) Merhavirus Merhavirus merida Merida virus (MERDV) Merhavirus tritaeniorhynchus Culex tritaeniorhynchus rhabdovirus (CTRV) Mousrhavirus Mousrhavirus moussa Moussa virus (MOUV) Ohlsrhavirus Ohlsrhavirus angeles Culex rhabdo-like virus Los Angeles (CRLVLA) Ohlsrhavirus culex Culex rhabdo-like virus (CRLV) 2875Taxonomic update of phylum Negarnaviricota 1 3 Table 4 (continued) Genus Species¶ Virus (Abbreviation)& Ohlsrhavirus lobeira Lobeira virus (LOBV) Ohlsrhavirus northcreek North Creek virus (NORCV) Ohlsrhavirus ohlsdorf Ohlsdorf virus (OHLDV) Ohlsrhavirus pseudovishnui Culex pseudovishnui rhabdo-like virus (CpRLV) Ohlsrhavirus riverside Riverside virus (RISV) Ohlsrhavirus tongilchon Tongilchon virus 1 (TCHV-1) Perhabdovirus Perhabdovirus anguilla eel virus American (EVA) eel virus European X (EVEX) Perhabdovirus leman Leman virus (LeRV) Perhabdovirus perca perch rhabdovirus (PRV) Perhabdovirus trutta lake trout rhabdovirus (LTRV) Replylivirus Replylivirus allogus anole lyssa-like virus 1 (ALLV-1) Sawgrhavirus Sawgrhavirus connecticut Connecticut virus (CNTV) Sawgrhavirus longisland Long Island tick rhabdovirus (LITRV) Sawgrhavirus minto New Minto virus (NMV) Sawgrhavirus sawgrass Sawgrass virus (SAWV) Scophrhavirus Scophrhavirus chanodychthys Wuhan redfin culter dimarhabdovirus (WhRFCRV) Scophrhavirus maximus Scophthalmus maximus rhabdovirus (SMRV) Sigmavirus Sigmavirus affinis Drosophila affinis sigmavirus (DAffSV) Sigmavirus ananassae Drosophila ananassae sigmavirus (DAnaSV) Sigmavirus capitata Ceratitis capitata sigmavirus (CCapSV) Sigmavirus domestica Wuhan fly virus 2 (WhFV-2) Sigmavirus hippoboscid Wuhan louse fly virus 9 (WhLFV-9) Sigmavirus hubei Hubei diptera virus 9 (HbDV-9) Sigmavirus immigrans Drosophila immigrans sigmavirus (DImmSV) Sigmavirus lousefly Wuhan louse fly virus 10 (WhLFV-10) Sigmavirus melanogaster Drosophila melanogaster sigmavirus (DMelSV) Sigmavirus muscina Muscina stabulans sigmavirus (MStaSV) Sigmavirus myga Hubei diptera virus 10 (HbDV-10) Sigmavirus obscura Drosophila obscura sigmavirus (DObsSV) Sigmavirus shayang Shayang fly virus 2 (SyFV-1) Sigmavirus sturtevanti Drosophila sturtevanti sigmavirus (DStuSV) Sigmavirus tristis Drosophila tristis sigmavirus (DTriSV) Sigmavirus wuhan Wuhan house fly virus 1 (WhHFV-1) Sigmavirus ying Hubei dimarhabdovirus 1 (HbDRV-1) Siniperhavirus Siniperhavirus chuatsi Chinese rice-field eel rhabdovirus (CrERV) hybrid snakehead rhabdovirus (HSHRV) Siniperca chuatsi rhabdovirus (SCRV) Micropterus salmoides rhabdovirus (MSRV) Siniperhavirus zoarces eelpout rhabdovirus (EPRV) Sprivivirus Sprivivirus cyprinus spring viremia of carp virus (SVCV) Sprivivirus esox grass carp rhabdovirus (GrCRV) pike fry rhabdovirus (PFRV) tench rhabdovirus (TenRV) Sripuvirus Sripuvirus almpiwar Almpiwar virus (ALMV) Sripuvirus chaco Chaco virus (CHOV) Sripuvirus charleville Charleville virus (CHVV) Sripuvirus cuiaba Cuiaba virus (CUIV) Sripuvirus hainan Hainan black-spectacled toad rhabdovirus (HnBSTRV) Sripuvirus madureira Sena Madureira virus (SMV) 2876 J. H. Kuhn et al. 1 3 Table 4 (continued) Genus Species¶ Virus (Abbreviation)& Sripuvirus niakha Niakha virus (NIAV) Sripuvirus sripur Sripur virus (SRIV) Sunrhavirus Sunrhavirus alexandria Burg el Arab virus (BEAV) Sunrhavirus bimbo Bimbo virus (BBOV) Sunrhavirus boteke Boteke virus (BOTV) Sunrhavirus dillard Dillard’s Draw virus (DDRV) Sunrhavirus garba Garba virus (GARV) Sunrhavirus harrison Harrison Dam virus (HARDV) Sunrhavirus kolongo Kolongo virus (KOLV) Sunrhavirus kwatta Kwatta virus (KWAV) Sunrhavirus matariya Matariya virus (MTYV) Sunrhavirus nasoule Nasoule virus (NASV) Sunrhavirus oakvale Oak Vale virus (OVV) Sunrhavirus ouango Ouango virus (OUAV) Sunrhavirus sandjimba Sandjimba virus (SJAV) Sunrhavirus sunguru Sunguru virus (SUNV) Sunrhavirus walkabout Walkabout Creek virus (WACV) Tibrovirus Tibrovirus alphaekpoma Ekpoma virus 1 (EKV-1) Tibrovirus beatrice Beatrice Hill virus (BHV) Tibrovirus betaekpoma Ekpoma virus 2 (EKV-2) Tibrovirus coastal Coastal Plains virus (CPV) Tibrovirus congo Bas-Congo virus (BASV) Tibrovirus sweetwater Sweetwater Branch virus (SWBV) Tibrovirus tibrogargan Bivens Arm virus (BAV) Tibrogargan virus (TIBV) Tupavirus Tupavirus durham Durham virus (DURV) Tupavirus klamath Klamath virus (KLAV) Tupavirus tupaia tupaia rhabdovirus (TUPV) Vesiculovirus Vesiculovirus alagoas vesicular stomatitis Alagoas virus (VSAV) Vesiculovirus bogdanovac Yug Bogdanovac virus (YBV) Vesiculovirus carajas Carajás virus (CJSV) Vesiculovirus chandipura Chandipura virus (CHPV) Vesiculovirus cocal Cocal virus (COCV) Vesiculovirus eptesicus American bat vesiculovirus (ABVV) Vesiculovirus indiana vesicular stomatitis Indiana virus (VSIV) Vesiculovirus isfahan Isfahan virus (ISFV) Vesiculovirus jurona Jurona virus (JURV) Vesiculovirus malpais Malpais Spring virus (MSPV) Vesiculovirus maraba Maraba virus (MARAV) Vesiculovirus mediterranean Mediterranean bat virus (MBV) Vesiculovirus morreton Morreton virus (MORV) Vesiculovirus newjersey vesicular stomatitis New Jersey virus (VSNJV) Vesiculovirus perinet Perinet virus (PERV) Vesiculovirus piry Piry virus (PIRYV) Vesiculovirus radi Radi virus (RADV) Vesiculovirus rhinolophus Jinghong bat virus (JhBV) Qiongzhong bat virus (QZBV) Vesiculovirus yinshui Yinshui bat virus (YSBV) Zarhavirus Zarhavirus zahedan Zahedan rhabdovirus (ZARV) 2877Taxonomic update of phylum Negarnaviricota 1 3 Table 4 (continued) Genus Species¶ Virus (Abbreviation)& Subfamily Betarhabdovirinae Alphanucleorhabdovirus Alphanucleorhabdovirus colocasiae taro vein chlorosis virus (TaVCV) Alphanucleorhabdovirus constrictae constricta yellow dwarf virus (CYDV) Alphanucleorhabdovirus joa joa yellow blotch associated virus (JYBaV) Alphanucleorhabdovirus maydis maize mosaic virus (MMV) Alphanucleorhabdovirus melongenae eggplant mottled dwarf virus (EMDV) Alphanucleorhabdovirus morogoromaydis Morogoro maize-associated virus (MMaV) Alphanucleorhabdovirus oryzae rice transitory yellowing virus (RTYV) rice yellow stunt virus (RYSV) Alphanucleorhabdovirus physostegiae Physostegia chlorotic mottle virus (PhCMoV) Alphanucleorhabdovirus pruni peach virus 1 (PeV1) Alphanucleorhabdovirus tritici wheat yellow striate virus (WYSV) Alphanucleorhabdovirus tuberosum potato yellow dwarf virus (PYDV) Alphanucleorhabdovirus zeairanense maize Iranian mosaic virus (MIMV) Betanucleorhabdovirus Betanucleorhabdovirus bacopae Bacopa monnieri virus 2 (BmV-2) Betanucleorhabdovirus cardamomi cardamom vein clearing virus (CdVCV) Betanucleorhabdovirus daturae datura yellow vein virus (DYVV) Betanucleorhabdovirus loti birds-foot trefoil-associated virus (BFTV) Betanucleorhabdovirus mali apple rootstock virus A (ApRVA) Betanucleorhabdovirus medicagonis alfalfa-associated nucleorhabdovirus (AaNV) Betanucleorhabdovirus retesonchi Sonchus yellow net virus (SYNV) Betanucleorhabdovirus ribes blackcurrant-associated rhabdovirus (BCaRV) Betanucleorhabdovirus venasonchi sowthistle yellow vein virus (SYVV) Betanucleorhabdovirus zanthoxyli Zhuye pepper nucleorhabdovirus (ZPNRV) Cytorhabdovirus Cytorhabdovirus alphatrifolii Trifolium pratense virus A (TpVA) Cytorhabdovirus alphawuhaninsectum Wuhan insect virus 4 (WuIV-4) Cytorhabdovirus bacopae Bacopa monnieri virus 1 (BmV-1) Cytorhabdovirus betatrifolii Trifolium pratense virus B (TpVB) Cytorhabdovirus betawuhaninsectum Wuhan insect virus 5 (WuIV-5) Cytorhabdovirus brassicae broccoli necrotic yellows virus (BNYV) Cytorhabdovirus brassicicolae cabbage cytorhabdovirus 1 (CCyV-1) Cytorhabdovirus broussonetiae paper mulberry mosaic associated virus (PMuMaV) Cytorhabdovirus caricae papaya virus E (PpVE) Cytorhabdovirus chrysanthemi chrysanthemum yellow dwarf associated virus (CYDaV) Cytorhabdovirus colocasiae Colocasia bobone disease-associated virus (CBDaV) Cytorhabdovirus cucurbitae cucurbit cytorhabdovirus 1 (CuCV-1) Cytorhabdovirus festucae Festuca leaf streak virus (FLSV) Cytorhabdovirus flaviyerbamate yerba mate chlorosis-associated virus (YmCaV) Cytorhabdovirus fragariae strawberry-associated virus 1 (SaV-1) strawberry virus 1 (StrV-1) Cytorhabdovirus fragariarugosus strawberry crinkle virus (SCV) Cytorhabdovirus gammawuhaninsectum Wuhan insect virus 6 (WuIV-6) Cytorhabdovirus gramineae northern cereal mosaic virus (NCMV) Cytorhabdovirus hordei barley yellow striate mosaic virus (BYSMV) Cytorhabdovirus lactucamaculante lettuce yellow mottle virus (LYMoV) Cytorhabdovirus lactucanecante lettuce necrotic yellows virus (LNYV) Cytorhabdovirus lycopersici tomato yellow mottle-associated virus (TYMaV) Cytorhabdovirus maydis maize-associated cytorhabdovirus (MaCV) Cytorhabdovirus maysflavostriatis maize yellow striate virus (MYSV) Cytorhabdovirus medicagonis alfalfa dwarf virus (ADV) 2878 J. H. Kuhn et al. 1 3 Table 4 (continued) Genus Species¶ Virus (Abbreviation)& Cytorhabdovirus oryzae rice stripe mosaic virus (RSMV) Cytorhabdovirus persimmon persimmon virus A (PeVA) Cytorhabdovirus rosae rose virus R (RVR) Cytorhabdovirus rubus raspberry vein chlorosis virus (RVCV) Cytorhabdovirus sonchi Sonchus virus (SonV) Cytorhabdovirus trichosanthei Trichosanthes associated rhabdovirus 1 (TrARV1) Cytorhabdovirus tritici wheat American striate mosaic virus (WASMV) Cytorhabdovirus yerbamate yerba mate virus A (YmVA) Dichorhavirus Dichorhavirus citri citrus chlorotic spot virus (CiCSV) Dichorhavirus clerodendri clerodendrum chlorotic spot virus (ClCSV) Dichorhavirus coffeae coffee ringspot virus (CoRSV) Dichorhavirus leprosis citrus leprosis virus N (CiLV-N) Dichorhavirus orchidaceae orchid fleck virus (OFV) Gammanucleorhabdovirus Gammanucleorhabdovirus maydis maize fine streak virus (MFSV) Varicosavirus Varicosavirus alopecuri Alopecurus myosuroides varicosavirus 1 (AMVV-1) Varicosavirus lactucae lettuce big-vein associated virus (LBVaV) Varicosavirus trifolii red clover associated varicosavirus (RCaVV) Subfamily Gammarhabdovirinae Novirhabdovirus Novirhabdovirus hirame hirame rhabdovirus (HIRRV = HIRV) Novirhabdovirus piscine viral hemorrhagic septicemia virus (VHSV) Novirhabdovirus salmonid infectious hematopoietic necrosis virus (IHNV) Novirhabdovirus snakehead snakehead rhabdovirus (SHRV) Unassigned (to subfamilies) Alphacrustrhavirus Alphacrustrhavirus wenling Wenling crustacean virus 10 (WlCV-10) Alphacrustrhavirus zhejiang Wenling crustacean virus 11 (WlCV-11) Alphadrosrhavirus Alphadrosrhavirus hubei Wuhan house fly virus 2 (WhHFV-2) Alphadrosrhavirus shayang Shayang fly virus 3 (SyFV-3) Alphahymrhavirus Alphahymrhavirus cinereus hymenopteran rhabdo-related virus 38 (HyRRV-38) Alphahymrhavirus hirtum hymenopteran rhabdo-related virus 109 (HyRRV-109) Alphahymrhavirus neglectus Lasius neglectus virus 2 (LnegV-2) Alphahymrhavirus radians hymenopteran rhabdo-related virus 46 (HyRRV-46) Betahymrhavirus Betahymrhavirus austriaca hymenopteran rhabdo-related virus 23 (HyRRV-23) Betahymrhavirus heterodontonyx hymenopteran rhabdo-related virus 24 (HyRRV-24) Betanemrhavirus Betanemrhavirus hubei Hubei rhabdo-like virus 9 (HbRLV-9) Betanemrhavirus shayang Shayang ascaridia galli virus 2 (SyAGV-2) Betapaprhavirus Betapaprhavirus frugiperda Spodoptera frugiperda rhabdovirus (SfruRV) Betapaprhavirus sylvina lepidopteran rhabdo-related virus 34 (LeRRV-34) Betaricinrhavirus Betaricinrhavirus chimay Chimay rhabdovirus (CRV) Betaricinrhavirus scapularis blacklegged tick rhabdovirus 1 (BLTRV-1) Family Sunviridae Sunshinevirus Sunshinevirus reptilis Sunshine Coast virus (SunCV) Family Xinmoviridae Alasvirus Alasvirus muscae Húběi diptera virus 11 (HbDV-11) Anphevirus Anphevirus xinchengense Xīnchéng mosquito virus (XcMV) Doupovirus Doupovirus australiaense Culex mononega-like virus 2 (CMLV-2) Draselvirus Draselvirus dentati Húběi rhabdo-like virus 7 (HbRLV-7) Drunivirus Drunivirus chambonense Drosophila unispina virus 1 (DuniV-1) Gambievirus Gambievirus bolahunense Bolahun virus (BLHV) 2879Taxonomic update of phylum Negarnaviricota 1 3 Table 4 (continued) Genus Species¶ Virus (Abbreviation)& Gambievirus senegalense Gambie virus (GAMV) Gylbovirus Gylbovirus aagae Aedes anphevirus (AeAV) Hoptevirus Hoptevirus orthopteris Húběi orthoptera virus 5 (HbOV-5) Madalivirus Madalivirus amapaense Anopheles marajoara virus (AnMV) Madalivirus amazonaense Anopheles darlingi virus (AnDV) Pelmivirus Pelmivirus eymattense hymenopteran anphe-related virus OKIAV71 (HARV-71) Triniovirus Triniovirus yonagoense Culex tritaeniorhynchus anphevirus (CtAV) Ulegvirus Ulegvirus freckenfeldense odonatan anphe-related virus OKIAV59 (OARV-59) Note that viruses are real objects that are assigned to concepts that are called taxa. Species, genera, subfamilies, families, and orders are taxa 1 Includes: Newcastle disease virus (NDV) ¶ Taxon names are always italicized and always begin with a capital letter & Virus names are not italicized and are not capitalized, except if the name or a name component is a proper noun. This column lists the virus names with their correct (lack of) capitalization. Lists of viruses within a given species are provisional at this point and will likely be amended in the near future Table 5 ICTV-accepted taxonomy of the order Goujianvirales (Negarnaviricota: Haploviricotina: Yunchangviricetes) as of March 2022 Note that viruses are real objects that are assigned to concepts that are called taxa. Species, genera, fami- lies, and orders are taxa ¶ Taxon names are always italicized and always begin with a capital letter & Virus names are not italicized and are not capitalized, except if the name or a name component is a proper noun. This column lists the virus names with their correct (lack of) capitalization Genus Species¶ Virus (Abbreviation)& Family Yueviridae Yuyuevirus Yuyuevirus beihaiense Běihǎi sesarmid crab virus 3 (BhSCV-3) Yuyuevirus shaheense Shāhé yuèvirus-like virus 1 (ShYLV-1) 2880 J. H. Kuhn et al. 1 3 Table 6 ICTV-accepted taxonomy of the order Bunyavirales (Negarnaviricota: Polyploviricotina: Ellioviricetes) as of March 2022 Genus Species¶ Virus (Abbreviation)& Family Arenaviridae Antennavirus Hairy antennavirus Wēnlǐng frogfish arenavirus 2 (WlFAV-2) Salmon antennavirus salmon pescarenavirus 1 (SPAV-1) salmon pescarenavirus 2 (SPAV-2) Striated antennavirus Wēnlǐng frogfish arenavirus 1 (WlFAV-1) Hartmanivirus Haartman hartmanivirus Haartman Institute snake virus 1 (HISV-1) Haartman Institute snake virus 2 (HISV-2) Heimat hartmanivirus andere Heimat virus 1 (aHeV-1) Muikkunen hartmanivirus Dante Muikkunen virus 1 (DaMV-1) Schoolhouse hartmanivirus old schoolhouse virus 1 (OScV-1) old schoolhouse virus 2 (OScV-2) Setpatvet hartmanivirus SetPatVet virus 1 (SPVV-1) Zurich hartmanivirus veterinary pathology Zurich virus 1 (VPZV-1) veterinary pathology Zurich virus 2 (VPZV-2) Mammarenavirus Allpahuayo mammarenavirus Allpahuayo virus (ALLV) Alxa mammarenavirus Alxa virus (ALXV) Argentinian mammarenavirus Junín virus (JUNV) Bear Canyon mammarenavirus Bear Canyon virus (BCNV) Brazilian mammarenavirus Sabiá virus (SBAV) Cali mammarenavirus Pichindé virus (PICHV) Chapare mammarenavirus Chapare virus (CHAPV) Chevrier mammarenavirus Lìjiāng virus (LIJV) Cupixi mammarenavirus Cupixi virus (CUPXV) Flexal mammarenavirus Flexal virus (FLEV) Gairo mammarenavirus Gairo virus (GAIV) Guanarito mammarenavirus Guanarito virus (GTOV) Ippy mammarenavirus Ippy virus (IPPYV) Kitale mammarenavirus Kitale virus (KTLV) Lassa mammarenavirus Lassa virus (LASV) Latino mammarenavirus Latino virus (LATV) Loei River mammarenavirus Loei River virus (LORV) Lujo mammarenavirus Lujo virus (LUJV) Luna mammarenavirus Luli virus (LULV) Luna virus (LUAV) Lunk mammarenavirus Lunk virus (LNKV) Lymphocytic choriomeningitis mammare- navirus Dandenong virus (DANV) lymphocytic choriomeningitis virus (LCMV) Machupo mammarenavirus Machupo virus (MACV) Mammarenavirus bituense Bitu virus (BITV) Mammarenavirus kwanzaense Kwanza virus (KWAV) Mariental mammarenavirus Mariental virus (MRLV) Merino Walk mammarenavirus Merino Walk virus (MRWV) Mobala mammarenavirus mobala virus (MOBV) Mopeia mammarenavirus Mopeia virus (MPOV) Morogoro virus (MORV) Okahandja mammarenavirus Okahandja virus (OKAV) Oliveros mammarenavirus Oliveros virus (OLVV) Paraguayan mammarenavirus Paraná virus (PRAV) Planalto mammarenavirus Aporé virus (APOV) 2881Taxonomic update of phylum Negarnaviricota 1 3 Table 6 (continued) Genus Species¶ Virus (Abbreviation)& Pirital mammarenavirus Pirital virus (PIRV) Ryukyu mammarenavirus Ryukyu virus (RYKV) Serra do Navio mammarenavirus Amaparí virus (AMAV) Solwezi mammarenavirus Solwezi virus (SOLV) Souris mammarenavirus souris virus (SOUV) Tacaribe mammarenavirus Tacaribe virus (TCRV) Tamiami mammarenavirus Tamiami virus (TMMV) Wenzhou mammarenavirus Wēnzhōu virus (WENV) Whitewater Arroyo mammarenavirus Big Brushy Tank virus (BBRTV) Catarina virus (CTNV) Skinner Tank virus (SKTV) Tonto Creek virus (TTCV) Whitewater Arroyo virus (WWAV) Xapuri mammarenavirus Xapuri virus (XAPV) Reptarenavirus California reptarenavirus CAS virus (CASV) Giessen reptarenavirus University of Giessen virus 1 (UGV-1) University of Giessen virus 2 (UGV-2) University of Giessen virus 3 (UGV-3) Golden reptarenavirus Golden Gate virus (GOGV) Ordinary reptarenavirus tavallinen suomalainen mies virus 2 (TSMV-2) Rotterdam reptarenavirus ROUT virus (ROUTV) University of Helsinki virus 1 (UHV-1) Family Cruliviridae Lincruvirus Lincruvirus europense European shore crab virus 1 (EscV-1) Lincruvirus sinense Chinese mitten crab virus 1 (CmcV-1) Lincruvirus wenlingense Wēnlǐng crustacean virus 9 (WlCV-9) Family Discoviridae Orthodiscovirus Orthodiscovirus coniellae Coniothyrium diplodiella negative-stranded RNA virus 1 (CdNSRV-1) Plasmopara viticola lesion associated mycobunyavirales-like virus 9 (PvLAM-LV-9) Orthodiscovirus hispaniae Plasmopara viticola lesion associated mycobunyavirales-like virus 8 (PvLAM-LV8) Orthodiscovirus iberiae Plasmopara viticola lesion associated mycobunyavirales-like virus 4 (PvLAM-LV4) Orthodiscovirus missouriense Penicillium discovirus (PDV) Orthodiscovirus penicillii Penicillium roseopurpureum negative ssRNA virus 1 (PrNssV-1) Family Fimoviridae Emaravirus Emaravirus aceris maple mottle-associated virus (MaMaV) Emaravirus actinidiae Actinidia chlorotic ringspot-associated virus (AcCRaV) Emaravirus cajani pigeonpea sterility mosaic virus 1 (PPSMV-1) Emaravirus camelliae Camellia japonica-associated virus 1 (CjaV-1) Emaravirus cercidis redbud yellow ringspot-associated virus (RYRaV) Emaravirus chrysanthemi chrysanthemum mosaic-associated virus (ChMaV) Emaravirus cordylinae ti ringspot-associated virus (TiRSaV) Emaravirus fici fig mosaic virus (FMV) Emaravirus idaeobati raspberry leaf blotch virus (RLBV) Emaravirus kiwii Actinidia virus 2 (AcV-2) 2882 J. H. Kuhn et al. 1 3 Table 6 (continued) Genus Species¶ Virus (Abbreviation)& Emaravirus parkinsoniae palo verde broom virus (PVBV) Emaravirus perillae perilla mosaic virus (PerMV) Emaravirus pistaciae Pistacia virus B (PiVB) Emaravirus populi aspen mosaic-associated virus (AsMaV) Emaravirus pyri pear chlorotic leaf spot-associated virus (PCLSaV) Emaravirus quercus common oak ringspot-associated virus (CORaV) Emaravirus rosae rose rosette virus (RRV) Emaravirus rubi blackberry leaf mottle-associated virus (BLMaV) Emaravirus sorbi European mountain ash ringspot-associated virus (EMARaV) Emaravirus syringae lilac chlorotic ringspot-associated virus (LiCRaV) Emaravirus toordali pigeonpea sterility mosaic virus 2 (PPSMV-2) Emaravirus tritici High Plains wheat mosaic virus (HPWMoV) Emaravirus verbanni Camellia japonica-associated virus 2 (CjaV-2) Emaravirus ziziphi jujube yellow mottle-associated virus (JYMaV) Family Hantaviridae Subfamily Actantavirinae Actinovirus Batfish actinovirus Wēnlǐng minipizza batfish virus (WEMBV) Goosefish actinovirus Wēnlǐng yellow goosefish virus (WEYGV) Perch actinovirus Bern perch virus (BRPV) Spikefish actinovirus Wēnlǐng red spikefish virus (WERSV) Subfamily Agantavirinae Agnathovirus Hagfish agnathovirus Wēnlǐng hagfish virus (WEHV) Subfamily Mammantavirinae Loanvirus Brno loanvirus Brno virus (BRNV) Longquan loanvirus Lóngquán virus (LQUV) Mobatvirus Laibin mobatvirus Láibīn virus (LAIV) Lena mobatvirus Lena virus (LENV) Nova mobatvirus Nova virus (NVAV) Quezon mobatvirus Quezon virus (QZNV) Xuan Son mobatvirus Xuân Sơn virus (XSV) Orthohantavirus Andes orthohantavirus Andes virus (ANDV) Castelo dos Sonhos virus (CASV) Lechiguanas virus (LECV = LECHV) Orán virus (ORNV) Asama orthohantavirus Asama virus (ASAV) Asikkala orthohantavirus Asikkala virus (ASIV) Bayou orthohantavirus bayou virus (BAYV) Catacamas virus (CATV) Black Creek Canal orthohantavirus Black Creek Canal virus (BCCV) Bowe orthohantavirus Bowé virus (BOWV) Bruges orthohantavirus Bruges virus (BRGV) Cano Delgadito orthohantavirus Caño Delgadito virus (CADV) Cao Bang orthohantavirus Cao Bằng virus (CBNV) Liánghé virus (LHEV) Choclo orthohantavirus Choclo virus (CHOV) Dabieshan orthohantavirus Dàbiéshān virus (DBSV) Dobrava-Belgrade orthohantavirus Dobrava virus (DOBV) Kurkino virus (KURV) Saaremaa virus (SAAV) 2883Taxonomic update of phylum Negarnaviricota 1 3 Table 6 (continued) Genus Species¶ Virus (Abbreviation)& Sochi virus (SOCV) El Moro Canyon orthohantavirus Carrizal virus (CARV) El Moro Canyon virus (ELMCV) Huitzilac virus (HUIV) Fugong orthohantavirus Fúgòng virus (FUGV) Fusong orthohantavirus Fǔsōng virus (FUSV) Hantaan orthohantavirus Amur virus (AMRV) Hantaan virus (HTNV) Soochong virus (SOOV) Jeju orthohantavirus Jeju virus (JJUV) Kenkeme orthohantavirus Kenkeme virus (KKMV) Khabarovsk orthohantavirus Khabarovsk virus (KHAV) Topografov virus (TOPV) Laguna Negra orthohantavirus Laguna Negra virus (LANV) Maripa virus (MARV) Rio Mamoré virus (RIOMV) Luxi orthohantavirus Lúxī virus (LUXV) Maporal orthohantavirus Maporal virus (MAPV) Montano orthohantavirus Montaño virus (MTNV) Necocli orthohantavirus Necoclí virus (NECV) Oxbow orthohantavirus Oxbow virus (OXBV) Prospect Hill orthohantavirus Prospect Hill virus (PHV) Puumala orthohantavirus Hokkaido virus (HOKV) Muju virus (MUJV) Puumala virus (PUUV) Robina orthohantavirus Robina virus (ROBV) Rockport orthohantavirus Rockport virus (RKPV) Sangassou orthohantavirus Sangassou virus (SANGV) Seewis orthohantavirus Seewis virus (SWSV) Seoul orthohantavirus gōu virus (GOUV) Seoul virus (SEOV) Sin Nombre orthohantavirus New York virus (NYV) Sin Nombre virus (SNV) Tatenale orthohantavirus Tatenale virus (TATV) Thailand orthohantavirus Anjozorobe virus (ANJZV) Serang virus (SERV) Thailand virus (THAIV) Tigray orthohantavirus Tigray virus (TIGV) Tula orthohantavirus Adler virus (ADLV) Tula virus (TULV) Yakeshi orthohantavirus Yákèshí virus (YKSV) Thottimvirus Imjin thottimvirus Imjin virus (MJNV) Thottapalayam thottimvirus Thottapalayam virus (TPMV) Subfamily Repantavirinae Reptillovirus Gecko reptillovirus Hǎinán oriental leaf-toed gecko virus (HOLGV) 2884 J. H. Kuhn et al. 1 3 Table 6 (continued) Genus Species¶ Virus (Abbreviation)& Family Leishbuviridae Shilevirus Shilevirus leptomonadis Leptomonas moramango virus (LEPMV) Family Mypoviridae Hubavirus Hubavirus myriapedis Húběi myriapoda virus 5 (HbMV-5) Family Nairoviridae Norwavirus Grotenhout norwavirus Grotenhout virus (GRHV) Ocetevirus Blattodean ocetevirus red goblin roach virus 1 (RGRV-1) Orthonairovirus Abu Hammad orthonairovirus Abū Ḥammād virus (AHV) Abu Mina orthonairovirus Abū Mīnā virus (AMV) Artashat orthonairovirus Artashat virus (ARTSV) Avalon orthonairovirus Avalon virus (AVAV) Chim orthonairovirus Chim virus (CHIMV) Bandia orthonairovirus Bandia virus (BDAV) Burana orthonairovirus Burana virus (BURV) Congoid orthonairovirus Aigai virus (AIGV) Crimean-Congo hemorrhagic fever ortho- nairovirus Crimean-Congo hemorrhagic fever virus (CCHFV) Dera Ghazi Khan orthonairovirus Dera Ghazi Khan virus (DGKV) Dugbe orthonairovirus Dugbe virus (DUGV) Erve orthonairovirus Erve virus (ERVEV) Estero Real orthonairovirus Estero Real virus (ERV) Gossas orthonairovirus Gossas virus (GOSV) Hazara orthonairovirus Hazara virus (HAZV) Huangpi orthonairovirus Huángpí tick virus 1 (HpTV-1) Hughes orthonairovirus Caspiy virus (CASV) Farallon virus (FARV) Great Saltee virus (GRSV) Hughes virus (HUGV) Raza virus (RAZAV) Issyk-kul orthonairovirus Issyk-kul virus (ISKV) Kasokero orthonairovirus Kasokero virus (KASV = KASOV) Keterah orthonairovirus Keterah virus (KTRV) Uzun-Agach virus (UZAV) Kupe orthonairovirus kupe virus (KUPEV) Leopards Hill orthonairovirus Leopards Hill virus (LPHV) Meram orthonairovirus Meram virus (MEMV) Nairobi sheep disease orthonairovirus Nairobi sheep disease virus (NSDV) Pacific Coast orthonairovirus Pacific Coast tick nairovirus (PCTNV) Punta orthonairovirus Punta Salinas virus (PSV) Qalyub orthonairovirus Geran virus (GERV) Qalyub virus (QYBV) Sakhalin orthonairovirus Sakhalin virus (SAKV) Tillamook virus (TILLV) Sapphire orthonairovirus Sapphire II virus (SAPV) Scot orthonairovirus Clo Mor virus (CMV = CLMV) Soldado orthonairovirus Soldado virus (SOLV) Tacheng orthonairovirus Tǎchéng tick virus 1 (TcTV-1) Taggert orthonairovirus Taggert virus (TAGV) 2885Taxonomic update of phylum Negarnaviricota 1 3 Table 6 (continued) Genus Species¶ Virus (Abbreviation)& Tamdy orthonairovirus Tamdy virus (TAMV) Thiafora orthonairovirus Thiafora virus (TFAV) Tofla orthonairovirus tofla virus (TFLV) Tunis orthonairovirus Tunis virus (TUNV) Vinegar Hill orthonairovirus Vinegar Hill virus (VINHV) Wenzhou orthonairovirus Wēnzhōu tick virus (WzTV) Yogue orthonairovirus Yogue virus (YOGV) Zirqa orthonairovirus Zirqa virus (ZIRV) Sabavirus South Bay sabavirus South Bay virus (SBV) Shaspivirus Spider shaspivirus Shāyáng spider virus 1 (SySV-1) Striwavirus Strider striwavirus Sānxiá water strider virus 1 (SxWSV-1) Xinspivirus Xinzhou xinspivirus Xīnzhōu spider virus (XSV) Family Peribunyaviridae Herbevirus Herbert herbevirus Herbert virus (HEBV) Kibale herbevirus Kibale virus (KIBV) Tai herbevirus Taï virus (TAIV) Khurdivirus Khurdivirus volgaense Khurdun virus (KURV) Lakivirus Lakivirus lakamhaense Lakamha virus (LAKV) Lambavirus Lambavirus wisconsinense largemouth bass bunyavirus (LBBV) Orthobunyavirus Abras orthobunyavirus Abras virus (ABRV) Acara orthobunyavirus Acará virus (ACAV) Moriche virus (MORV) Aino orthobunyavirus Aino virus (AINOV) Akabane orthobunyavirus Akabane virus (AKAV) Tinaroo virus (TINV) Yaba-7 virus (Y7V) Alajuela orthobunyavirus Alajuela virus (ALJV) San Juan virus (SJV) Anadyr orthobunyavirus Anadyr virus (ANADV) Ananindeua orthobunyavirus Ananindeua virus (ANUV) Anhembi orthobunyavirus Anhembi virus (AMBV) Anopheles A orthobunyavirus Anopheles A virus (ANAV) Arumateua virus (ARTV = ARMTV) Caraipé virus (CPEV = CRPV) Las Maloyas virus (LMV) Lukuni virus (LUKV) Trombetas virus (TRMV) Tucuruí virus (TUCV = TUCRV) Anopheles B orthobunyavirus Anopheles B virus (ANBV) Boracéia virus (BORV) Apeu orthobunyavirus Apeú virus (APEUV) Bakau orthobunyavirus Bakau virus (BAKV) Ketapang virus (KETV) Nola virus (NOLAV) Tanjong Rabok virus (TRV) Telok Forest virus (TFV) Batai orthobunyavirus Batai virus (BATV) Batama orthobunyavirus Batama virus (BMAV) Bellavista orthobunyavirus Bellavista virus (BELLV) 2886 J. H. Kuhn et al. 1 3 Table 6 (continued) Genus Species¶ Virus (Abbreviation)& Benevides orthobunyavirus Benevides virus (BVSV = BENV) Bertioga orthobunyavirus Bertioga virus (BERV) Cananéia virus (CNAV) Guaratuba virus (GTBV) Itimirim virus (ITIV) Mirim virus (MIRV) Bimiti orthobunyavirus bimiti virus (BIMV) Birao orthobunyavirus Birao virus (BIRV) Botambi orthobunyavirus Botambi virus (BOTV) Bozo orthobunyavirus Bozo virus (BOZOV) Brazoran orthobunyavirus brazoran virus (BRAZV) Bruconha orthobunyavirus Bruconha virus (BRUV) Buffalo Creek orthobunyavirus Buffalo Creek virus (BUCV) Bunyamwera orthobunyavirus Bunyamwera virus (BUNV) Germiston virus (GERV) Lokern virus (LOKV) Mboké virus (MBOV) Ngari virus (NRIV) Northway virus (NORV) Santa Rosa virus (SARV) Shokwe virus (SHOV) Stanfield virus (STAV) Xingu virus (XINV) Bushbush orthobunyavirus Benfica virus (BENV = BNFV) Bushbush virus (BSBV) Juan Díaz virus (JDV) Buttonwillow orthobunyavirus Buttonwillow virus (BUTV) Bwamba orthobunyavirus Bwamba virus (BWAV) Pongola virus (PGAV) Cache Valley orthobunyavirus Cache Valley virus (CVV) Cholul virus (CHLV) Tlacotalpan virus (TLAV) Cachoeira Porteira orthobunyavirus Cachoeira Porteira virus (CPOV) California encephalitis orthobunyavirus California encephalitis virus (CEV) Morro Bay virus (MBV) Capim orthobunyavirus Capim virus (CAPV) Caraparu orthobunyavirus Caraparú virus (CARV) El Huayo virus (EHUV) Itaya virus (ITYV) Ossa virus (OSSAV) Vinces virus (VINV) Cat Que orthobunyavirus Cát Quế virus (CQV) Oya virus (OYAV) Catu orthobunyavirus Catú virus (CATUV) Enseada orthobunyavirus Enseada virus (ENSV) Faceys paddock orthobunyavirus Facey’s paddock virus (FPV) Fort Sherman orthobunyavirus Fort Sherman virus (FSV) Gamboa orthobunyavirus Brus Laguna virus (BLAV) Calchaquí virus (CQIV) Gamboa virus (GAMV) 2887Taxonomic update of phylum Negarnaviricota 1 3 Table 6 (continued) Genus Species¶ Virus (Abbreviation)& Pueblo Viejo virus (PVV) Soberanía virus (SOBV) Gan Gan orthobunyavirus Gan Gan virus (GGV) Guajara orthobunyavirus Guajará virus (GJAV) Guama orthobunyavirus Guamá virus (GMAV) Guaroa orthobunyavirus Guaroa virus (GROV) Iaco orthobunyavirus Iaco virus (IACOV) Ilesha orthobunyavirus Ilesha virus (ILEV) Ingwavuma orthobunyavirus Ingwavuma virus (INGV) Jamestown Canyon orthobunyavirus Inkoo virus (INKV) Jamestown Canyon virus (JCV) Jerry Slough virus (JSV) South River virus (SORV) Jatobal orthobunyavirus Jatobal virus (JATV) Kaeng Khoi orthobunyavirus Kaeng Khoi virus (KKV) Kairi orthobunyavirus kairi virus (KRIV) Keystone orthobunyavirus Keystone virus (KEYV) Koongol orthobunyavirus koongol virus (KOOV) wongal virus (WONV) La Crosse orthobunyavirus La Crosse virus (LACV) Leanyer orthobunyavirus Leanyer virus (LEAV) Lumbo orthobunyavirus Lumbo virus (LUMV) Macaua orthobunyavirus Macauã virus (MCAV) Madrid orthobunyavirus Madrid virus (MADV) Maguari orthobunyavirus Maguari virus (MAGV) Playas virus (PLAV) Mahogany Hammock orthobunyavirus Mahogany Hammock virus (MHV) Main Drain orthobunyavirus Main Drain virus (MDV) Manzanilla orthobunyavirus Manzanilla virus (MANV) Inini virus (INIV) Mapputta orthobunyavirus Mapputta virus (MAPV) Maprik orthobunyavirus Maprik virus (MPKV) Marituba orthobunyavirus Gumbo Limbo virus (GLV) Marituba virus (MTBV) Murutucú virus (MURV) Nepuyo virus (NEPV) Restan virus (RESV) Zungarococha virus (ZUNV) Matruh orthobunyavirus Matruh virus (MTRV) Melao orthobunyavirus Melao virus (MELV) Moju orthobunyavirus Moju virus (MOJUV) Mermet orthobunyavirus Mermet virus (MERV) Minatitlan orthobunyavirus Minatitlán virus (MNTV) Palestina virus (PLSV) MPoko orthobunyavirus M'Poko virus (MPOV) Yaba-1 virus (Y1V) Nyando orthobunyavirus Eretmapodites virus (ERETV) Mojuí dos Campos virus (MDCV) Nyando virus (NDV) Olifantsvlei orthobunyavirus Bobia virus (BIAV) 2888 J. H. Kuhn et al. 1 3 Table 6 (continued) Genus Species¶ Virus (Abbreviation)& Dabakala virus (DABV) Olifantsvlei virus (OLIV) Oubi virus (OUBIV) Oriboca orthobunyavirus Itaquí virus (ITQV) Oriboca virus (ORIV) Oropouche orthobunyavirus Iquitos virus (IQTV) Madre de Dios virus (MDDV) Oropouche virus (OROV) Perdões virus (PDEV) Pintupo virus (PINTV) Oyo orthobunyavirus Oyo virus (OYOV) Patois orthobunyavirus Babahoya virus (BABV) Pahayokee virus (PAHV) Patois virus (PATV) Peaton orthobunyavirus peaton virus (PEAV) Potosi orthobunyavirus Potosi virus (POTV) Sabo orthobunyavirus Sabo virus (SABOV) San Angelo orthobunyavirus San Angelo virus (SAV) Sango orthobunyavirus Sango virus (SANV) Schmallenberg orthobunyavirus Douglas virus (DOUV) Sathuperi virus (SATV) Schmallenberg virus (SBV) Shamonda virus (SHAV) Sedlec orthobunyavirus Sedlec virus (SEDV) Serra do Navio orthobunyavirus Serra do Navio virus (SDNV) Shark River orthobunyavirus Shark River virus (SRV) Shuni orthobunyavirus Kaikalur virus (KAIV) Shuni virus (SHUV) Simbu orthobunyavirus Para virus (PARAV) Simbu virus (SIMV) Snowshoe hare orthobunyavirus Khatanga virus (KHATV) snowshoe hare virus (SSHV) Sororoca orthobunyavirus Sororoca virus (SORV) Tacaiuma orthobunyavirus CoAr 1071 virus (CA1071V) CoAr 3627 virus (CA3626V) Tacaiuma virus (TCMV) Virgin River virus (VRV) Tahyna orthobunyavirus Ťahyňa virus (TAHV) Tataguine orthobunyavirus Tataguine virus (TATV) Tensaw orthobunyavirus Tensaw virus (TENV) Tete orthobunyavirus Bahig virus (BAHV) Tete virus (TETEV) Tsuruse virus (TSUV) Weldona virus (WELV) Thimiri orthobunyavirus Thimiri virus (THIV) Timboteua orthobunyavirus Timboteua virus (TBTV) Triniti orthobunyavirus Triniti virus (TNTV) Trivittatus orthobunyavirus Achiote virus (ACHOV) Trivittatus virus (TVTV) Turlock orthobunyavirus Lednice virus (LEDV) 2889Taxonomic update of phylum Negarnaviricota 1 3 Table 6 (continued) Genus Species¶ Virus (Abbreviation)& Turlock virus (TURV) Umbre virus (UMBV) Utinga orthobunyavirus Utinga virus (UTIV) Witwatersrand orthobunyavirus Witwatersrand virus (WITV) Wolkberg orthobunyavirus Wolkberg virus (WBV) Wyeomyia orthobunyavirus Rio Pracupi virus Taiassui virus (TAIAV) Tucunduba virus (TUCV) Wyeomyia virus (WYOV) Pacuvirus Caimito pacuvirus Caimito virus (CAIV) Chilibre pacuvirus Chilibre virus (CHIV) Pacui pacuvirus Pacui virus (PACV) Rio Preto da Eva pacuvirus Rio Preto da Eva virus (RPEV) Tapirape pacuvirus Tapirapé virus (TAPV) Shangavirus Insect shangavirus Shuāngào insect virus 1 (SgIV-1) Family Phasmaviridae Feravirus Ferak feravirus Ferak virus (FRKV) Hemipteran feravirus hemipteran phasma-related virus OKIAV247 (HeFV) Neuropteran feravirus neuropteran phasma-related virus OKIAV248 (NeFV) Hymovirus Hymenopteran hymovirus 1 hymenopteran phasma-related virus OKIAV252 (HyHV-1) Hymenopteran hymovirus 2 hymenopteran phasma-related virus OKIAV250 (HyHV-2) Jonvirus Jonchet jonvirus jonchet virus (JONV) Orthophasmavirus Anopheles orthophasmavirus Anopheles triannulatus orthophasmavirus (AtOPV) Coleopteran orthophasmavirus coleopteran phasma-related virus OKIAV235 (CPRV) Culex orthophasmavirus Culex phasma-like virus (CPLV) Ganda orthophasmavirus Ganda bee virus (GBEEV) Hymenopteran orthophasmavirus 1 hymenopteran phasma-related virus OKIAV228 (HyOV-1) Hymenopteran orthophasmavirus 2 hymenopteran phasma-related virus OKIAV227 (HyOV-2) Kigluaik phantom orthophasmavirus Kigluaik phantom virus (KIGV) Niukluk phantom orthophasmavirus Niukluk phantom virus (NUKV) Odonate orthophasmavirus Húběi odonate virus 8 (HbOV-8) Orthophasmavirus barstukasense Barstukas virus (BARV) Orthophasmavirus coredoense Coredo virus (CORV) Orthophasmavirus flenense Flen virus (FLNV) Orthophasmavirus miglotasense Miglotas virus (MIGV) Qingling orthophasmavirus Húběi odonate virus 9 (HbOV-9) Wuchang cockroach orthophasmavirus 1 Wǔchāng cockroach virus 1 (WcCV-1) Wuhan mosquito orthophasmavirus 1 Wǔhàn mosquito virus 1 (WhMV-1) Wuhan mosquito orthophasmavirus 2 Wǔhàn mosquito virus 2 (WhMV-2) Sawastrivirus Sanxia sawastrivirus Sānxiá water strider virus 2 (SxWSV-2) Wuhivirus Insect wuhivirus Wǔhàn insect virus 2 (WhIV-2) Family Phenuiviridae Bandavirus Bhanja bandavirus Bhanja virus (BHAV) Dabie bandavirus severe fever with thrombocytopenia syndrome virus (SFTSV) Guertu bandavirus Gùěrtú virus (GTV) Heartland bandavirus Heartland virus (HRTV) Hunter Island bandavirus Albatross Island virus (ABIV) 2890 J. H. Kuhn et al. 1 3 Table 6 (continued) Genus Species¶ Virus (Abbreviation)& Kismaayo bandavirus Kismaayo virus (KISV) Lone star bandavirus lone star virus (LSV) Razdan bandavirus Razdan virus (RAZV) Beidivirus Dipteran beidivirus Húběi diptera virus 3 (HbDV-3) Coguvirus Citrus coguvirus citrus concave gum-associated virus (CCGaV) Coguvirus eburi citrus virus A (CiV-A) Grapevine coguvirus grapevine associated cogu-like virus 1 (GaCLV-1) Entovirus Entoleuca entovirus Entoleuca phenui-like virus 1 (EnPLV-1) Goukovirus Cumuto goukovirus Cumuto virus (CUMV) Gouleako goukovirus Gouléako virus (GOLV) Yichang insect goukovirus Yíchāng insect virus (YcIV) Horwuvirus Horsefly horwuvirus Wǔhàn horsefly virus (WhHV) Kimberley horwuvirus Fitzroy Crossing tenui-like virus 1 (FCTenV1) Hudivirus Dipteran hudivirus Húběi diptera virus 4 (HbDV-4) Hudovirus Lepidopteran hudovirus Húběi lepidoptera virus 1 (HbLV-1) Ixovirus Blackleg ixovirus blacklegged tick virus 1 (BLTV-1) Norway ixovirus Fairhair virus (FHAV) Scapularis ixovirus blacklegged tick virus 3 (BLTV-3) Laulavirus Grapevine laulavirus 2 grapevine associated cogu-like virus 2 (GaCLV-2) Grapevine laulavirus 3 grapevine associated cogu-like virus 3 (GaCLV-3) Grapevine laulavirus 4 grapevine associated cogu-like virus 4 (GaCLV-4) Laurel Lake laulavirus Laurel Lake virus (LLV) Lentinuvirus Lentinula lentinuvirus Lentinula edodes negative-strand RNA virus 2 (LeNSRV-2) Mobuvirus Mothra mobuvirus Mothra virus (MTHV) Narangue mobuvirus Ñarangue virus (NRGV) Phasivirus Badu phasivirus Badu virus (BADUV) Dipteran phasivirus Húběi diptera virus 5 (HbDV-5) Fly phasivirus Wǔhàn fly virus 1 (WhFV-1) Guadeloupe phasivirus Guadeloupe mosquito phasivirus (GMPV) Kimberley phasivirus Parry’s Creek phasivirus 1 (PCPhasV1) Phasi Charoen-like phasivirus Phasi Charoen-like virus (PCLV) Wutai mosquito phasivirus Wǔtái mosquito virus (WtMV) Phlebovirus Adana phlebovirus Adana virus (ADAV) Aguacate phlebovirus Aguacate virus (AGUV) Alcube phlebovirus Alcube virus (ACBV) Alenquer phlebovirus Alenquer virus (ALEV) Ambe phlebovirus Ambé virus (ABEV) Anhanga phlebovirus Anhangá virus (ANHV) Arumowot phlebovirus Arumowot virus (AMTV) Buenaventura phlebovirus Buenaventura virus (BUEV) Bogoria phlebovirus Bogoria virus (BGRV) Bujaru phlebovirus Bujaru virus (BUJV) Cacao phlebovirus Cacao virus (CACV) Campana phlebovirus Campana virus (CMAV) Candiru phlebovirus Ariquemes virus (ARQV) Candirú virus (CDUV) Jacundá virus (JCNV) Morumbi virus (MR(M)BV) Mucura virus (MCRV/MRAV) Serra Norte virus (SRNV) 2891Taxonomic update of phylum Negarnaviricota 1 3 Table 6 (continued) Genus Species¶ Virus (Abbreviation)& Chagres phlebovirus Chagres virus (CHGV) Cocle phlebovirus Coclé virus (CCLV) Corfou phlebovirus Corfou virus (CFUV) Dashli phlebovirus Dāshlī virus (DASV) Durania phlebovirus Durania virus (DRNV) Echarate phlebovirus Echarate virus (ECHV) Embossos phlebovirus Embossos virus (EMBV) Gabek phlebovirus Gabek forest virus (GFV) Gordil phlebovirus Gordil virus (GORV) Icoaraci phlebovirus Icoaraci virus (ICOV) Itaituba phlebovirus Itaituba virus (ITAV) Itaporanga phlebovirus Itaporanga virus (ITPV) Ixcanal phlebovirus Ixcanal virus (IXCV) Karimabad phlebovirus Karimabad virus (KARV) Kiborgoch phlebovirus Kiborgoch virus (KBGV) La Gloria phlebovirus La Gloria virus (LAGV) Lara phlebovirus Rio Claro virus (RICV) Leticia phlebovirus Leticia virus (LTCV) Maldonado phlebovirus Maldonado virus (MLOV) Massilia phlebovirus Massilia virus (MASV) Medjerda phlebovirus Medjerda Valley virus (MVV) Mona Grita phlebovirus Mona Grita virus (MOGV) Mukawa phlebovirus Mukawa virus (MKWV) Munguba phlebovirus Munguba virus (MUNV) Naples phlebovirus Arrábida virus (ARRV) Balkan virus (BALKV) Fermo virus (FERV) Granada virus (GRV = GRAV) Saddaguia virus (SADV) sandfly fever Naples virus (SFNV) Nique phlebovirus Nique virus (NIQV) Ntepes phlebovirus Ntepes virus (NTPV) Odrenisrou phlebovirus Odrénisrou virus (ODRV) Oriximina phlebovirus Oriximiná virus (ORXV) Pena Blanca phlebovirus Peña Blanca virus (PEBV) Penshurt phlebovirus Penshurt virus (PEHV) Perkerra phlebovirus Perkerra virus (PKEV) Punique phlebovirus Punique virus (PUNV) Punta Toro phlebovirus Capira virus (CAPIV) Punta Toro virus (PTV) Rift Valley fever phlebovirus Rift Valley fever virus (RVFV) Rio Grande phlebovirus Rio Grande virus (RGV) Saint Floris phlebovirus Saint-Floris virus (SAFV) Salanga phlebovirus Salanga virus (SLGV) Salehabad phlebovirus Adria virus (ADRV) Arbia virus (ARBV) Bregalaka virus (BREV) Olbia virus (OLBV) Salehabad virus (SALV) Zaba virus (ZABAV) 2892 J. H. Kuhn et al. 1 3 Table 6 (continued) Genus Species¶ Virus (Abbreviation)& Salobo phlebovirus Salobo virus (SLBOV) Sicilian phlebovirus sandfly fever Sicilian virus (SFSV) Tapara phlebovirus Tapará virus (TPRV) Tehran phlebovirus Tehran virus (THEV) Tico phlebovirus Tico virus (TICV) Toros phlebovirus Toros virus (TORV) Toscana phlebovirus Toscana virus (TOSV) Tres Almendras phlebovirus Tres Almendras virus (TRAV) Turuna phlebovirus Turuna virus (TUAV) Uriurana phlebovirus Uriurana virus (URIV) Urucuri phlebovirus Urucuri virus (URUV) Viola phlebovirus viola virus (VIOV) Zerdali phlebovirus Zerdali virus (ZERV) Pidchovirus Coleopteran pidchovirus coleopteran phenui-related virus 308 (CoPrV-308) Pidgey pidchovirus Pidgey virus (PGYV) Rubodvirus Apple rubodvirus 1 apple rubbery wood virus 1 (ARWV-1) Apple rubodvirus 2 apple rubbery wood virus 2 (ARWV-2) Grapevine rubodvirus 1 grapevine Garan dmak virus (GGDV) Grapevine rubodvirus 2 grapevine Muscat rose virus (GMRV) Tanzavirus Human tanzavirus Dar es Salaam virus (DeSV) Tenuivirus Echinochloa hoja blanca tenuivirus Echinochloa hoja blanca virus (EHBV) European wheat striate mosaic tenuivirus European wheat striate mosaic virus (EWSMV) Iranian wheat stripe tenuivirus Iranian wheat stripe virus (IWSV) Maize stripe tenuivirus maize stripe virus (MStV = MSpV) Melon tenuivirus melon chlorotic spot virus (MeCSV) Rice grassy stunt tenuivirus rice grassy stunt virus (RGSV) Rice hoja blanca tenuivirus rice hoja blanca virus (RHBV) Rice stripe tenuivirus rice stripe virus (RSV = RStV) Urochloa hoja blanca tenuivirus Urochloa hoja blanca virus (UHBV) Uukuvirus American dog uukuvirus American dog tick virus (ADAV) Dabieshan uukuvirus Dàbiéshān tick virus (DBSH) Grand Arbaud uukuvirus Grand Arbaud virus (GAV) Huangpi uukuvirus Huángpí tick virus 2 (HpTV-2) Kabuto mountain uukuvirus Kabuto mountain virus (KAMV) Kaisodi uukuvirus Kaisodi virus (KASDV) Lihan uukuvirus Lǐhán tick virus (LITV) Murre uukuvirus murre virus (MURV) Pacific coast uukuvirus Pacific coast tick virus (PACV) Precarious Point uukuvirus Precarious Point virus (PPV) Rukutama uukuvirus Rukutama virus (RUKV) Schmid uukuvirus Nile Warbler virus (NIWV) Silverwater uukuvirus Silverwater virus (SILV) Tacheng uukuvirus Tǎchéng tick virus 2 (TCGV) Uukuniemi uukuvirus Chizé virus (CHZV) Fin V 707 virus (FINV) Oceanside virus (OCV = OCEV) Pontevès virus (PTVV) St. Abbs Head virus (SAHV) Uukuniemi virus (UUKV) 2893Taxonomic update of phylum Negarnaviricota 1 3 Note that viruses are real objects that are assigned to concepts that are called taxa. Species, genera, subfamilies, families, and orders are taxa ¶ Taxon names are always italicized and always begin with a capital letter & Virus names are not italicized and are not capitalized, except if the name or a name component is a proper noun. This column lists the virus names with their correct (lack of) capitalization. Lists of viruses within a given species are provisional at this point and will likely be amended in the near future Genus Species¶ Virus (Abbreviation)& Yongjia uukuvirus Yǒngjiā tick virus (YONV) Zaliv Terpeniya uukuvirus Zaliv Terpeniya virus (ZTV) Wenrivirus Shrimp wenrivirus Mourilyan virus (MoV) Family Tospoviridae Orthotospovirus Alstroemeria necrotic streak orthotospovirus Alstroemeria necrotic streak virus (ANSV) Alstroemeria yellow spot orthotospovirus Alstroemeria yellow spot virus (AYSV) Bean necrotic mosaic orthotospovirus bean necrotic mosaic virus (BeNMV) Calla lily chlorotic spot orthotospovirus calla lily chlorotic spot virus (CCSV) Capsicum chlorosis orthotospovirus Capsicum chlorosis virus (CaCV) Chrysanthemum stem necrosis orthotospovirus Chrysanthemum stem necrosis virus (CSNV) Groundnut bud necrosis orthotospovirus groundnut bud necrosis virus (GBNV) Groundnut chlorotic fan spot orthotospovirus groundnut chlorotic fan-spot virus (GCFSV) Groundnut ringspot orthotospovirus groundnut ringspot virus (GRSV) Groundnut yellow spot orthotospovirus groundnut yellow spot virus (GYSV) Hippeastrum chlorotic ringspot orthotospovirus Hippeastrum chlorotic spot virus (HCRV) Impatiens necrotic spot orthotospovirus impatiens necrotic spot virus (INSV) Iris yellow spot orthotospovirus iris yellow spot virus (IYSV) Melon severe mosaic orthotospovirus melon severe mosaic virus (MSMV) Melon yellow spot orthotospovirus melon yellow spot virus (MYSV) Mulberry vein banding associated orthotospovirus mulberry vein banding-associated virus (MVBaV) Pepper chlorotic spot orthotospovirus pepper chlorotic spot virus (PCSV) Polygonum ringspot orthotospovirus Polygonum ringspot virus (PolRSV) Soybean vein necrosis orthotospovirus soybean vein necrosis virus (SVNV) Tomato chlorotic spot orthotospovirus tomato chlorotic spot virus (TCSV) Tomato spotted wilt orthotospovirus tomato spotted wilt virus (TSWV) Tomato yellow ring orthotospovirus tomato yellow ring virus (TYRV) Tomato zonate spot orthotospovirus tomato zonate spot virus (TZSV) Watermelon bud necrosis orthotospovirus watermelon bud necrosis virus (WBNV) Watermelon silver mottle orthotospovirus watermelon silver mottle virus (WSMoV) Zucchini lethal chlorosis orthotospovirus zucchini lethal chlorosis virus (ZLCV) Family Tulasviridae Orthotulasvirus Orthotulasvirus tulasnellae Tulasnella bunyavirales-like virus 1 (TB-LV) Family Wupedeviridae Wumivirus Wumivirus millepedae Wǔhàn millipede virus 2 (WhMV-2) Table 6 (continued) 2894 J. H. Kuhn et al. 1 3 Supplementary Information The online version contains supplemen- tary material available at https:// doi. org/ 10. 1007/ s00705- 022- 05546-z. Acknowledgements Members of the 2020–2023 International Com- mittee on Taxonomy of Viruses (ICTV): Piet Maes; Marco Marklewitz; Gustavo Palacios; Takahide Sasaya (笹谷孝英); Massimo Turina; Yong-Zhen Zhang (张永振)); Filoviridae (Jens H. Kuhn, Nadine Biedenkopf; Alexander Bukreyev; Kartik Chandran; Nicholas Di Paola; Pierre B. H. Formenty; Anthony Griffiths; Adam J. Hume; Elke Mühlberger; Sergey V. Netesov (Heтёcoв Cepгeй Bиктopoвич); Gustavo Palacios; Janusz T. Pawęska; Sophie Smither; Ayato Takada (高田礼人); Victoria Wahl); Fimoviridae (Michele Digiaro; Toufic Elbeaino; Kenji Kubota (久保田健嗣); Francisco M. Ochoa-Corona; Susanne von Bargen); Hantaviridae (Steven B. Bradfute; Charles H. Calisher; Boris Klempa; Jonas Klingström; Lies Laenen; Piet Maes; Nicole D. Tischler); Jingchuvirales (Jens H. Kuhn; Nolwenn M. Dheilly; Nicholas Di Paola; Sandra Junglen; Sofia Paraskevopoulou (Σοφία Παρασκευοπούλου); Mang Shi (施莽)); Lispiviridae (Jun-Min Li (李俊敏); Sofia Paraskevopoulou (Σοφία Παρασκευοπούλου); Gongyin Ye (叶恭银)); Monjiviricetes (Jens H. Kuhn; Nicholas Di Paola; Ralf G. Dietzgen; W. Paul Duprex; Juliana Freitas-Astúa; Timothy H. Hyndman; Dàohóng Jiāng (姜道宏); Piet Maes; Sofia Paraskevopoulou (Σοφία Παρασκευοπούλου); Janusz T. Pawęska; Dennis Rubbenstroth; Peter J. Walker; Yong-Zhen Zhang (张永振)); Table 7 ICTV-accepted taxonomy of the order Articulavirales (Negarnaviricota: Polyploviricotina: Insthoviricetes) as of March 2022 Note that viruses are real objects that are assigned to concepts that are called taxa. Species, genera, fami- lies, and orders are taxa ¶ Taxon names are always italicized and always begin with a capital letter & Virus names are not italicized and are not capitalized, except if the name or a name component is a proper noun. This column lists the virus names with their correct (lack of) capitalization Genus Species¶ Virus (Abbreviation)& Family Amnoonviridae Tilapinevirus Tilapinevirus tilapiae tilapia lake virus (TiLV) Family Orthomyxoviridae Alphainfluenzavirus Alphainfluenzavirus influenzae influenza A virus (FLUAV) Betainfluenzavirus Betainfluenzavirus influenzae influenza B virus (FLUBV) Deltainfluenzavirus Deltainfluenzavirus influenzae influenza D virus (FLUDV) Gammainfluenzavirus Gammainfluenzavirus influenzae influenza C virus (FLUCV) Isavirus Isavirus salaris infectious salmon anemia virus (ISAV) Mykissvirus Mykissvirus tructae rainbow trout orthomyxovirus (RbtOV) steelhead trout orthomyxovirus (SttOV-1) Quaranjavirus Quaranjavirus araguariense Araguari virus (ARAV) Quaranjavirus chadense Lake Chad virus (LKCV) Quaranjavirus johnstonense Johnston Atoll virus (JAV) Quaranjavirus quaranfilense Quaranfil virus (QRFV) Quaranjavirus tyulekense Tyulek (Tjuloc) virus (TLKV) Quaranjavirus wellfleetense Wellfleet Bay virus (WFBV) Sardinovirus Sardinovirus pilchardi pilchard orthomyxovirus (POMV) Thogotovirus Thogotovirus bourbonense Bourbon virus (BRBV) Thogotovirus dhoriense Dhori virus (DHOV) Thogotovirus josense Jos virus (JOSV) Thogotovirus ozense Oz virus (OZV) Thogotovirus sinuense Sinu virus (SINUV) Thogotovirus thailandense Thailand tick thogotovirus (TT-THOV) Thogotovirus thogotoense Thogoto virus (THOV) Thogotovirus upoluense Upolu virus (UPOV) Arenaviridae (Jens H. Kuhn; Michael J. Buchmeier; Rémi N. Charrel; Juan Carlos de la Torre; Xavier de Lamballerie; Jean-Paul J. Gonzalez; Stephan Günther; Jussi Hepojoki; Igor S. Lukashevich; Sheli R. Radoshitzky; Víctor Romanowski; Manuela Sironi; Mark D. Stenglein); Artoviridae (Jens H. Kuhn; Ralf G. Dietzgen; Nikos Vasilakis (Νίκος Βασιλάκης); Gongyin Ye (叶恭银); Arnfinn Lodden Økland); Aspiviridae (Elena Dal Bó; Selma Gago-Zachert; María Laura García; Tomohide Natsuaki (夏秋知英); José A. Navarro; Yutaro Neriya (煉谷裕太朗); John Hammond; Vicente Pallás; Carina A. Reyes; Takahide Sasaya (笹谷孝英); Ioannis E. Tzanetakis; Anna Maria Vaira; Martin Verbeek); Bornaviridae (Jens H. Kuhn; Thomas Briese; Ralf Dürrwald; Mas- ayuki Horie (堀江真行); Timothy H. Hyndman; Norbert Nowotny; Florian Pfaff; Dennis Rubbenstroth; Keizō Tomonaga (朝長啓造); Ellioviricetes (Jens H. Kuhn, Scott Adkins; Juan Carlos de la Torre; Michele Digiaro; Holly R. Hughes; Sandra Junglen; Amy J. Lambert; 2895Taxonomic update of phylum Negarnaviricota 1 3 Mymonaviridae (María A. Ayllón; Dàohóng Jiāng (姜道宏); Hideki Kondō (近藤秀樹); Shin-Yi L. Marzano; Massimo Turina); Nairoviridae (Jens H. Kuhn, Sergey V. Alkhovsky (Aльxoвcкий Cepгeй Bлaдимиpoвич), Tatjana Avšič-Županc, Éric Bergeron; Felic- ity Burt; Koray Ergünay; Aura R. Garrison; Marco Marklewitz; Ali Mirazimi; Gustavo Palacios; Anna Papa (Άννα Παπά); Janusz T. Pawęska; Jessica R. Spengler); Negarnaviricota (Jens H. Kuhn; Eugene V. Koonin; Mart Krupovic; Yuri I. Wolf); Nyamiviridae (Jens H. Kuhn; Ralf G. Dietzgen; Andrew E. Firth; Sofia Paraskevopoulou (Σοφία Παρασκευοπούλου); Nikos Vasilakis (Νίκος Βασιλάκης)); Orthomyxoviridae (Justin Bahl; Inmaculada Casas; Adolfo García- Sastre; Seiji Hongō (本郷誠治); Sergio H. Marshall; John W. McCau- ley; Gabriele Neumann; Colin R. Parrish; Daniel R. Pérez; Jonathan A. Runstadler; Martin Schwemmle); Paramyxoviridae (Anne Balkema-Buschmann; J. Felix Drexler; W. Paul Duprex; Piet Maes; Richard K. Plemper; Bert Vanmechelen); Peribunyaviridae (Sergey V. Alkhovsky (Aльxoвcкий Cepгeй Bлaдимиpoвич); Martin Beer; Carol D. Blair; Charles H. Calisher; William M. de Souza; Holly R. Hughes; Amy J. Lambert; Marco Marklewitz); Phasmaviridae (Matthew J. Ballinger; Michael A. Drebot; Sandra Junglen; Alex Pauvolid-Corrêa); Phenuiviridae (Thomas Briese; Francesco Di Serio; Martin H. Gro- schup; Gilda B. Jonson; Yutaro Neriya (煉谷裕太朗; Gustavo Pala- cios; Takahide Sasaya (笹谷孝英); Jin-Won Song (송진원)); Yasuhiro Tomitaka (冨髙保弘)); Pneumoviridae (Paul A. Brown; Ursula J. Buchholz; Rik L. de Swart; J. Felix Drexler; W. Paul Duprex; Andrew J. Easton; Jiànróng Lǐ (李建荣); Kirsten Spann; Natalie J. Thornburg; Bernadette van den Hoogen; John V. Williams); Rhabdoviridae (Nicolas Bejerman; Kim R. Blasdell; Ralf G. Dietz- gen; Anthony R. Fooks; Juliana Freitas-Astúa; Hideki Kondō (近藤秀 樹); Ivan V. Kuzmin; Pedro L. Ramos-González; Mang Shi (施莽); Robert B. Tesh; Noël Tordo; Nikos Vasilakis (Νίκος Βασιλάκης); Peter J. Walker; Anna E. Whitfield); Sunviridae (Jens H. Kuhn; Timothy H. Hyndman); Tospoviridae (Scott Adkins; Holly R. Hughes; Rayapati Naidu; Renato O. Resende; Massimo Turina; Anna E. Whitfield); Xinmoviridae (Guozhong Feng (冯国忠); Sofia Paraskevopoulou (Σοφία Παρασκευοπούλου); Stephen Sharpe) Study Groups. 2020–2023 ICTV Chair of the Animal dsRNA and ssRNA- Viruses Subcommittee (Jens H. Kuhn); 2020–2023 ICTV Chair of the Archaeal Viruses Subcommittee (Mart Krupovic); 2020–2023 ICTV Chair of the Plant Viruses Subcommittee (Luisa Rubino); 2020–2023 ICTV Chair Fungal and Protist Viruses Subcommittee (Sead Sabanadzovic); and ICTV Proposal Secretary (Peter J. Walker) Chair of the 2020–2023 ICTV Ellioviricetes, Filoviridae, and Negarnaviricota Study Groups (Jens H. Kuhn); Chair of the 2020– 2023 ICTV Tospoviridae Study Group (Scott Adkins); Chair of the 2020–2023 ICTV Arenaviridae Study Group (Juan Carlos de la Torre); Chair of the 2020–2023 ICTV Jingchuvirales Study Group (Nicholas Di Paola); Chair of the 2020–2023 ICTV Artoviridae and Nyamiviri- dae Study Groups (Ralf G. Dietzgen); Chair of the 2020–2023 ICTV Fimoviridae Study Group (Michele Digiaro); Chair of the 2020–2023 ICTV Pneumoviridae Study Group (W. Paul Duprex); Co-Chair of the 2020–2023 ICTV Rhabdoviridae Study Group (Juliana Freitas- Astúa; Peter J. Walker); Chair of the 2020–2023 ICTV Orthomyxo- viridae Study Group (Adolfo García-Sastre); Chair of the 2020–2023 ICTV Peribunyaviridae Study Group (Holly R. Hughes); Chair of the 2020–2023 ICTV Sunviridae Study Group (Timothy H. Hyndman); Chair of the 2020–2023 ICTV Mymonaviridae Study Group (Dàohóng Jiāng (姜道宏)); Chair of the 2020–2023 ICTV Phasmaviridae Study Group (Sandra Junglen); Chair of the 2020–2023 ICTV Hantaviridae, Monjiviricetes, and Paramyxoviridae Study Groups (Piet Maes); Chair of the 2020–2023 ICTV Nairoviridae Study Group (Gustavo Palacios); Co-Chair of the 2020–2023 ICTV Phenuiviridae Study Group (Gus- tavo Palacios; Takahide Sasaya (笹谷孝英)); Resigned Chair of the 2020–2023 ICTV Paramyxoviridae Study Group (Richard K. Plemper); Chair of the 2020–2023 ICTV Lispiviridae and Xinmoviridae Study Groups (Sofia Paraskevopoulou (Σοφία Παρασκευοπούλου)); Chair of the 2020–2023 ICTV Bornaviridae Study Group (Dennis Rubben- stroth); and Chair of the 2020–2023 ICTV Aspiviridae Study Group (Anna Maria Vaira). Funding This work was supported in part through Laulima Govern- ment Solutions, LLC prime contract with the US National Institute of Allergy and Infectious Diseases (NIAID) under Contract No. HHSN272201800013C. J.H.K. performed this work as an employee of Tunnell Government Services (TGS), a subcontractor of Laulima Gov- ernment Solutions, LLC under Contract No. HHSN272201800013C. This work was also funded in part by Contract No. HSHQDC- 15-C-00064 awarded by DHS S&T for the management and operation of The National Biodefense Analysis and Countermeasures Center, a federally funded research and development center operated by the Battelle National Biodefense Institute (V.W.); and NIH contract HHSN272201000040I/HHSN27200004/D04 and grant R24AI120942 (N.V., R.B.T.). S.S. acknowledges support from the Mississippi Agri- cultural and Forestry Experiment Station (MAFES), USDA-ARS pro- ject 58-6066-9-033 and the National Institute of Food and Agriculture, US Department of Agriculture, Hatch Project, under Accession Num- ber 1021494. Declarations Conflict of interest The authors declare no conflicts of interest. Ethical approval This article does not contain any studies with human participants or animals performed by any of the authors. Compliance with ethical standards The views and conclusions con- tained in this document are those of the authors and should not be interpreted as necessarily representing the official policies, either ex- pressed or implied, of the US Department of the Army, the US De- partment of Defense, the US Department of Health and Human Ser- vices, the US Department of Homeland Security (DHS) Science and Technology Directorate (S&T), or of the institutions and companies affiliated with the authors. 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Kuhn kuhnjens@mail.nih.gov Scott Adkins scott.adkins@ars.usda.gov Sergey V. Alkhovsky (Aльxoвcкий Cepгeй Bлaдимиpoвич) salkh@yandex.ru Tatjana Avšič-Županc Tatjana.Avsic@mf.uni-lj.si María A. Ayllón mariaangeles.ayllon@upm.es Justin Bahl justin.bahl@uga.edu 2900 J. H. Kuhn et al. 1 3 Anne Balkema-Buschmann anne.balkema-buschmann@fli.de Matthew J. Ballinger ballinger@biology.msstate.edu Martina Bandte martina.bandte@agrar.hu-berlin.de Martin Beer martin.beer@fli.de Nicolas Bejerman bejerman.nicolas@inta.gob.ar Éric Bergeron exj8@cdc.gov Nadine Biedenkopf nadine.biedenkopf@staff.uni-marburg.de Laurent Bigarré laurent.bigarre@anses.fr Carol D. Blair carol.blair@colostate.edu Kim R. Blasdell kim.blasdell@csiro.au Steven B. Bradfute sbradfute@salud.unm.edu Thomas Briese thomas.briese@columbia.edu Paul A. Brown paul.brown@anses.fr Rémy Bruggmann remy.bruggmann@bioinformatics.unibe.ch Ursula J. Buchholz ubuchholz@niaid.nih.gov Michael J. Buchmeier m.buchmeier@uci.edu Alexander Bukreyev alexander.bukreyev@utmb.edu Felicity Burt burtfj@ufs.ac.za Carmen Büttner carmen.buettner@agrar.hu-berlin.de Charles H. Calisher calisher6920@gmail.com Thierry Candresse thierry.candresse@inrae.fr Jeremy Carson jeremy.carson@bioconsulting.com.au Inmaculada Casas icasas@isciii.es Kartik Chandran kartik.chandran@einstein.yu.edu Rémi N. Charrel remi.charrel@univ-amu.fr Yuya Chiaki (千秋祐也) chiakiy151@affrc.go.jp Anya Crane anya.crane@nih.gov Mark Crane Mark.Crane@csiro.au Laurent Dacheux laurent.dacheux@pasteur.fr Elena Dal Bó elenadalbocastanion@gmail.com Juan Carlos de la Torre juanct@scripps.edu Xavier de Lamballerie xavier.de-lamballerie@univ-amu.fr William M. de Souza wmarciel@hotmail.com Rik L. de Swart rik.deswart@wur.nl Nolwenn M. Dheilly nolwenn.dheilly@anses.fr Nicholas Di Paola nicholas.dipaola.civ@health.mil Francesco Di Serio francesco.diserio@ipsp.cnr.it Ralf G. Dietzgen r.dietzgen@uq.edu.au Michele Digiaro digiaro@iamb.it J. Felix Drexler felix.drexler@charite.de W. Paul Duprex pduprex@pitt.edu Ralf Dürrwald duerrwaldr@rki.de Andrew J. Easton A.J.Easton@warwick.ac.uk Toufic Elbeaino elbeaino@iamb.it Koray Ergünay ekoray@hacettepe.edu.tr Guozhong Feng (冯国忠) fengguozhong@caas.cn Claudette Feuvrier cfeuvrier@jura.fr Andrew E. Firth aef24@cam.ac.uk Anthony R. Fooks Tony.Fooks@apha.gov.uk Pierre B. H. Formenty formentyp@who.int Juliana Freitas-Astúa juliana.astua@embrapa.br Selma Gago-Zachert selma.gago-zachert@bct.uni-halle.de 2901Taxonomic update of phylum Negarnaviricota 1 3 María Laura García marialauragarcia2010@gmail.com Adolfo García-Sastre Adolfo.Garcia-Sastre@mssm.edu Aura R. Garrison aura.r.garrison.civ@health.mil Scott E. Godwin Scott.Godwin@nre.tas.gov.au Jean-Paul J. Gonzalez Jean.Paul.Gonzalez@georgetown.edu Joëlle Goüy de Bellocq joellegouy@gmail.com Anthony Griffiths ahgriff@bu.edu Martin H. Groschup Martin.Groschup@fli.de Stephan Günther guenther@bni.uni-hamburg.de John Hammond John.Hammond@usda.gov Jussi Hepojoki jussi.hepojoki@helsinki.fi; jussi.hepojoki@uzh.ch Melanie M. Hierweger melanie_hierweger@web.de Seiji Hongō (本郷誠治) shongou@med.id.yamagata-u.ac.jp Masayuki Horie (堀江真行) mhorie@omu.ac.jp Hidenori Horikawa (堀川英則) hidenori.horikawa@pref.aichi.lg.jp Holly R. Hughes ltr8@cdc.gov Adam J. Hume hume@bu.edu Timothy H. Hyndman T.Hyndman@murdoch.edu.au Dàohóng Jiāng (姜道宏) daohongjiang@mail.hzau.edu.cn Gilda B. Jonson g.jonson@irri.org Sandra Junglen sandra.junglen@charite.de Fujio Kadono (上遠野冨士夫) acaritick@yahoo.co.jp David G. Karlin davidgkarlin@gmail.com Boris Klempa boris.klempa@savba.sk Jonas Klingström jonas.klingstrom@ki.se Michel C. Koch michel.koch@vetsuisse.unibe.ch Hideki Kondō (近藤秀樹) hkondo@okayama-u.ac.jp Eugene V. Koonin koonin@ncbi.nlm.nih.gov Jarmila Krásová krasova@ivb.cz Mart Krupovic mart.krupovic@pasteur.fr Kenji Kubota (久保田健嗣) kubotak@affrc.go.jp Ivan V. Kuzmin ivankuzmin1965@gmail.com Lies Laenen lies.laenen@uzleuven.be Amy J. Lambert ahk7@CDC.GOV Jiànróng Lǐ (李建荣) li.926@osu.edu Jun-Min Li (李俊敏) lijunmin@nbu.edu.cn François Lieffrig f.lieffrig@cergroupe.be Igor S. Lukashevich isluka01@louisville.edu Dongsheng Luo dongshengluo@outlook.com Piet Maes piet.maes@kuleuven.be Marco Marklewitz marco.marklewitz@finddx.org Sergio H. Marshall sergio.marshall@pucv.cl Shin-Yi L. Marzano shinyi.marzano@usda.gov John W. McCauley john.mccauley@crick.ac.uk Ali Mirazimi Ali.Mirazimi@ki.se Peter G. Mohr Peter.Mohr@csiro.au Nick J. G. Moody Nick.Moody@csiro.au Yasuaki Morita (森田泰彰) ojigotakeguro@piano.ocn.ne.jp Richard N. Morrison Richard.Morrison@nre.tas.gov.au Elke Mühlberger muehlber@bu.edu Rayapati Naidu naidu.rayapati@wsu.edu Tomohide Natsuaki (夏秋知英) natsuaki@cc.utsunomiya-u.ac.jp 2902 J. H. Kuhn et al. 1 3 José A. Navarro janavarr@ibmcp.upv.es Yutaro Neriya (煉谷裕太朗) neriya@a.utsunomiya-u.ac.jp Sergey V. Netesov (Heтёcoв Cepгeй Bиктopoвич) netesov.s@nsu.ru Gabriele Neumann gabriele.neumann@wisc.edu Norbert Nowotny Norbert.Nowotny@vetmeduni.ac.at Francisco M. Ochoa-Corona ochoaco@okstate.edu Gustavo Palacios gustavo.palacios@mssm.edu Laurane Pallandre laurane.pallandre@anses.fr Vicente Pallás vpallas@ibmcp.upv.es Anna Papa (Άννα Παπά) annap.med@gmail.com Sofia Paraskevopoulou (Σοφία Παρασκευοπούλου) paraskevopoulous@rki.de Colin R. Parrish crp3@cornell.edu Alex Pauvolid-Corrêa pauvolid-correa@ufv.br Janusz T. Pawęska januszp@nicd.ac.za Daniel R. Pérez dperez1@uga.edu Florian Pfaff Florian.Pfaff@fli.de Richard K. Plemper rplemper@gsu.edu Thomas S. Postler tp2405@cumc.columbia.edu Françoise Pozet fpozet@jura.fr Sheli R. Radoshitzky sheliijo@gmail.com Pedro L. Ramos-González plrg1970@gmail.com Marius Rehanek marius.rehanek@agrar.hu-berlin.de Renato O. Resende rresende@unb.br Carina A. Reyes carinar@biol.unlp.edu.ar Víctor Romanowski vromanowski@gmail.com Dennis Rubbenstroth Dennis.Rubbenstroth@fli.de Luisa Rubino luisa.rubino@ipsp.cnr.it Artemis Rumbou (Άρτεμις Ρούμπου) artemis.rumbou@agrar.hu-berlin.de Jonathan A. Runstadler jonathan.runstadler@tufts.edu Melanie Rupp rupp@fishdoc.ch Sead Sabanadzovic SSabanadzovic@entomology.msstate.edu Takahide Sasaya (笹谷孝英) tsasaya@affrc.go.jp Heike Schmidt-Posthaus heike.schmidt@unibe.ch Martin Schwemmle martin.schwemmle@uniklinik-freiburg.de Torsten Seuberlich torsten.seuberlich@vetsuisse.unibe.ch Stephen R. Sharpe Stephen.Sharpe@westernsydney.edu.au Mang Shi (施莽) shim23@mail.sysu.edu.cn Manuela Sironi manuela.sironi@BP.LNF.it Sophie Smither sjsmither@mail.dstl.gov.uk Jin-Won Song (송진원) jwsong@korea.ac.kr Kirsten M. Spann Kirsten.Spann@qut.edu.au Jessica R. Spengler JSpengler@cdc.gov Mark D. Stenglein Mark.Stenglein@ColoState.edu Ayato Takada (高田礼人) atakada@czc.hokudai.ac.jp Robert B. Tesh rtesh@utmb.edu Jana Těšíková jana.tesikova@gmail.com Natalie J. Thornburg nax3@cdc.gov Nicole D. Tischler ntischler@cienciavida.org Yasuhiro Tomitaka (冨髙保弘) yasut@affrc.go.jp Keizō Tomonaga (朝長啓造) tomonaga.keizo.5r@kyoto-u.ac.jp; tomonaga@infront.kyoto-u.ac.jp Noël Tordo ntordo@pasteur.fr Kenta Tsunekawa kenta_tsunekawa@pref.aichi.lg.jp 2903Taxonomic update of phylum Negarnaviricota 1 3 Massimo Turina massimo.turina@ipsp.cnr.it Ioannis E. Tzanetakis itzaneta@uark.edu Anna Maria Vaira annamaria.vaira@ipsp.cnr.it Bernadette van den Hoogen b.vandenhoogen@erasmusmc.nl Bert Vanmechelen bert.vanmechelen@kuleuven.be Nikos Vasilakis (Νίκος Βασιλάκης) nivasila@utmb.edu Martin Verbeek martin.verbeek@wur.nl Susanne von Bargen susanne.von.bargen@agrar.hu-berlin.de Jiro Wada (和田治郎) wadaj@niaid.nih.gov Victoria Wahl victoria.wahl@st.dhs.gov Peter J. Walker peter.walker@uq.edu.au Anna E. Whitfield awhitfi@ncsu.edu John V. Williams jvw@chp.edu Yuri I. Wolf wolf@ncbi.nlm.nih.gov Junki Yamasaki (山﨑淳紀) junki_yamasaki@ken3.pref.kochi.lg.jp Hironobu Yanagisawa (栁澤広宣) hironobu_yanagisa090@maff.go.jp Gongyin Ye (叶恭银) chu@zju.edu.cn Yong-Zhen Zhang (张永振) zhangyongzhen@icdc.cn Arnfinn Lodden Økland arnfinn.lodden.okland@zoetis.com 1 Integrated Research Facility at Fort Detrick (IRF-Frederick), National Institute of Allergy and Infectious Diseases, National Institutes of Health (NIH), Fort Detrick, Frederick, MD, USA 2 United States Department of Agriculture, Agricultural Research Service, US Horticultural Research Laboratory, Fort Pierce, FL, USA 3 D.I. Ivanovsky Institute of Virology of N.F. Gamaleya National Center on Epidemiology and Microbiology of Ministry of Health of Russian Federation, Moscow, Russia 4 Institute of Microbiology and Immunology, Faculty of Medicine, University of Ljubljana, Ljubljana, Slovenia 5 Centro de Biotecnología y Genómica de Plantas, Universidad Politécnica de Madrid—Instituto Nacional de Investigación y Tecnología Agraria y Alimentaria, Campus de Montegancedo, Pozuelo de Alarcón, Madrid, Spain 6 Departamento de Biotecnología-Biología Vegetal, Escuela Técnica Superior de Ingeniería Agronómica, Alimentaria y de Biosistemas, Universidad Politécnica de Madrid, Madrid, Spain 7 Center for Ecology of Infectious Diseases, Department of Infectious Diseases, Department of Epidemiology and Biostatistics, Insitute of Bioinformatics, University of Georgia, Athens, GA, USA 8 Friedrich-Loeffler-Institut, Institute of Novel and Emerging Infectious Diseases, Greifswald-Insel Riems, Greifswald, Germany 9 Department of Biological Sciences, Mississippi State University, Mississippi State, Starkville, MS, USA 10 Division Phytomedicine, Faculty of Life Sciences, Humboldt-Universität zu Berlin, Berlin, Germany 11 Institute of Diagnostic Virology, Friedrich-Loeffler-Institut, Greifswald-Insel Riems, Germany 12 UFyMA, INTA-CONICET, Córdoba, Argentina 13 Division of High-Consequence Pathogens and Pathology, Viral Special Pathogens Branch, Centers for Disease Control and Prevention, Atlanta, GA, USA 14 Institute of Virology, Philipps-University Marburg, Marburg, Germany 15 French Agency for Food, Environmental and Occupational Heath Safety ANSES, Laboratory of Ploufragan-Plouzané-Niort, Ploufragan, France 16 Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, CO, USA 17 Commonwealth Scientific and Industrial Research Organisation (CSIRO), Australian Centre for Disease Preparedness, Geelong, VIC, Australia 18 University of New Mexico Health Sciences Center, Albuquerque, NM, USA 19 Center for Infection and Immunity, and Department of Epidemiology, Mailman School of Public Health, Columbia University, New York, New York, USA 20 French Agency for Food, Environmental and Occupational Heath Safety ANSES, Laboratory of Ploufragan-Plouzané-Niort, Ploufragan, France 21 Interfaculty Bioinformatics Unit, University of Bern, Bern, Switzerland 22 RNA Viruses Section, Laboratory of Infectious Diseases, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Bethesda, MD, USA 23 Department of Molecular Biology and Biochemistry, University of California, Irvine, CA, USA 24 Galveston National Laboratory, The University of Texas Medical Branch at Galveston, Galveston, TX, USA 25 Division of Virology, National Health Laboratory Service and Division of Virology, University of the Free State, Bloemfontein, Republic of South Africa 26 Colorado State University, Fort Collins, CO, USA 27 Univ. Bordeaux, INRAE, UMR BFP, Villenave d’Ornon, France 28 Centre for Aquatic Animal Health and Vaccines, Department of Natural Resources and Environment Tasmania, Launceston, TAS, Australia 2904 J. H. Kuhn et al. 1 3 29 Respiratory Virus and Influenza Unit, National Microbiology Center, Instituto de Salud Carlos III, Madrid, Spain 30 Department of Microbiology and Immunology, Albert Einstein College of Medicine, Bronx, NY, USA 31 Unité des Virus Emergents (Aix-Marseille Univ-IRD 190-Inserm 1207-IHU Méditerranée Infection), Marseille, France 32 Division of Fruit Tree and Tea Pest Control Research, Institute for Plant Protection, NARO, Tsukuba, Ibaraki, Japan 33 Integrated Research Facility at Fort Detrick, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Frederick, MD, USA 34 CSIRO Australian Centre for Disease Preparedness, East Geelong, VIC, Australia 35 Institut Pasteur, Université Paris Cité, Unit Lyssavirus Epidemiology and Neuropathology, National Reference Center for Rabies, WHO Collaborating Center for Reference and Research on Rabies, Paris, France 36 CIDEFI. Facultad de Ciencias Agrarias y Forestales, Universidad de La Plata, La Plata, Argentina 37 Department of Immunology and Microbiology IMM-6, The Scripps Research Institute, La Jolla, CA, USA 38 Unité des Virus Emergents (Aix-Marseille Univ-IRD 190-Inserm 1207-IHU Méditerranée Infection), Marseille, France 39 World Reference Center for Emerging Viruses and Arboviruses and Department of Microbiology and Immunology, The University of Texas Medical Branch at Galveston, Galveston, TX, USA 40 Department of Virology, Wageningen Bioveterinary Research, Lelystad, The Netherlands 41 UMR 1161 Virology ANSES/INRAE/ENVA, ANSES Animal Health Laboratory, Maisons-Alfort, France 42 United States Army Medical Research Institute of Infectious Diseases, Fort Detrick, Frederick, MD, USA 43 Istituto per la Protezione Sostenibile delle Piante, Consiglio Nazionale delle Ricerche, Bari, Italy 44 Queensland Alliance for Agriculture and Food Innovation, The University of Queensland, St. Lucia, QLD, Australia 45 CIHEAM, Istituto Agronomico Mediterraneo di Bari, Valenzano, Italy 46 Institute of Virology, Charité-Universitätsmedizin Berlin, corporate member of Freie Universität Berlin, Humboldt-Universität zu Berlin, Berlin, Germany 47 School of Medicine, University of Pittsburgh, Pittsburgh, PA, USA 48 Robert Koch Institut, Berlin, Germany 49 School of Life Sciences, University of Warwick, Coventry, UK 50 Department of Medical Microbiology, Virology Unit, Hacettepe University Faculty of Medicine, Ankara, Turkey 51 Walter Reed Biosystematics Unit (WRBU), Smithsonian Institution, Museum Support Center, Suitland, MD, USA 52 One Health Branch, Walter Reed Army Institute of Research (WRAIR), Silver Spring, MD, USA 53 Department of Entomology, Smithsonian Institution– National Museum of Natural History (NMNH), Washington, DC, USA 54 China National Rice Research Institute, Hangzhou, China 55 Departemental Analysis Laboratory, Poligny, France 56 Department of Pathology, University of Cambridge, Cambridge, UK 57 Animal and Plant Health Agency, Weybridge, Surrey, UK 58 World Health Organization, Geneva, Switzerland 59 Embrapa Cassava and Fruits, Cruz das Almas, Bahia, Brazil 60 Institute of Biochemistry and Biotechnology, Martin Luther University Halle-Wittenberg, Halle/Saale, Germany 61 Instituto de Biotecnología y Biología Molecular, Facultad de Ciencias Exactas, CONICET UNLP, La Plata, Argentina 62 Icahn School of Medicine at Mount Sinai, New York, NY, USA 63 United States Army Medical Research Institute of Infectious Diseases, Fort Detrick, Frederick, MD, USA 64 Department of Microbiology and Immunology, Division of Biomedical Graduate Research Organization, School of Medicine, Georgetown University, Washington, DC, USA 65 Institute of Vertebrate Biology of the Czech Academy of Sciences, Brno, Czech Republic 66 Department of Microbiology and National Emerging Infectious Diseases Laboratories, Boston University School of Medicine, Boston, MA, USA 67 Institute of Novel and Emerging Infectious Diseases, Friedrich-Loeffler-Institut, Greifswald-Insel Riems, Germany 68 Department of Virology, WHO Collaborating Centre for Arboviruses and Hemorrhagic Fever Reference and Research, Bernhard-Nocht Institute for Tropical Medicine, Hamburg, Germany 69 United States Department of Agriculture, Agricultural Research Service, USNA, Floral and Nursery Plants Research Unit, Beltsville, MD, USA 70 Department of Virology, University of Helsinki, Medicum, Helsinki, Finland 71 Institute of Veterinary Pathology, Vetsuisse Faculty, University of Zurich, Zurich, Switzerland 72 Division of Neurological Sciences, Vetsuisse Faculty, University of Bern, Bern, Switzerland 73 Department of Infectious Diseases, Yamagata University Faculty of Medicine, Yamagata, Japan 74 Graduate School of Veterinary Science, Osaka Metropolitan University, Izumisano, Osaka, Japan 75 Osaka International Research Center for Infectious Diseases, Osaka Metropolitan University, Izumisano, Osaka, Japan 76 Aichi Agricultural Research Center, Nagakute, Aichi, Japan 77 Centers for Disease Control and Prevention, Fort Collins, CO, USA 2905Taxonomic update of phylum Negarnaviricota 1 3 78 Department of Microbiology and National Emerging Infectious Diseases Laboratories, Boston University School of Medicine, Boston, MA, USA 79 Center for Emerging Infectious Diseases Policy and Research, Boston University School of Medicine, Boston, Massachusetts, USA 80 School of Veterinary Medicine, Murdoch University, Murdoch, WA, Australia 81 State Key Laboratory of Agricultural Microbiology, Huazhong Agricultural University, Wuhan, Hubei Province, China 82 International Rice Research Institute, College, Los Baños, 4032 Laguna, Philippines 83 Clinical Plant Science Center, Hosei University, Tokyo, Japan 84 Institute of Virology, Biomedical Research Center, Slovak Academy of Sciences, Bratislava, Slovakia 85 Department of Medicine Huddinge, Center for Infectious Medicine, Karolinska Institutet, Karolinska University Hospital, Stockholm, Sweden 86 Division of Neurological Sciences, Vetsuisse Faculty, University of Bern, Bern, Switzerland 87 Institute of Plant Science and Resources, Okayama University, Kurashiki, Japan 88 National Center for Biotechnology Information, National Library of Medicine, National Institutes of Health, Bethesda, MD, USA 89 Institut Pasteur, Université Paris Cité, CNRS UMR6047, Archaeal Virology Unit, Paris, France 90 Institute for Plant Protection, NARO, Tsukuba, Ibaraki, Japan 91 The University of Texas Medical Branch at Galveston, Galveston, TX, USA 92 KU Leuven, Rega Institute, Zoonotic Infectious Diseases unit, Leuven, Belgium 93 Department of Laboratory Medicine, University Hospitals Leuven, Leuven, Belgium 94 Centers for Disease Control and Prevention, Fort Collins, CO, USA 95 Department of Veterinary Biosciences, College of Veterinary Medicine, The Ohio State University, Columbus, OH, USA 96 Institute of Plant Virology, Ningbo University, Ningbo, China 97 CER Groupe, Fish Disease Laboratory, Marche-En-Famenne, Belgium 98 Department of Pharmacology and Toxicology, School of Medicine, and the Center for Predictive Medicine for Biodefense and Emerging Infectious Diseases, University of Louisville, Louisville, KY, USA 99 Institut Pasteur, Université Paris Cité, Unit Lyssavirus Epidemiology and Neuropathology, Paris, France 100 CAS Key Laboratory of Special Pathogens and Biosafety, Wuhan Institute of Virology, Chinese Academy of Sciences, Wuhan, Hubei, China 101 KU Leuven, Rega Institute, Zoonotic Infectious Diseases unit, Leuven, Belgium 102 FIND, the global alliance for diagnostics, Geneva, Switzerland 103 Instituto de Biología-Laboratorio de Genética Molecular-Campus Curauma, Valparaíso, Chile 104 United States Department of Agriculture, Agricultural Research Service, Toledo, OH, USA 105 Worldwide Influenza Centre, Francis Crick Institute, London, UK 106 Folkhalsomyndigheten, Stockholm, Sweden 107 Kochi Agricultural Research Center, Kochi, Japan 108 Department of Microbiology and National Emerging Infectious Diseases Laboratories, Boston University School of Medicine, Boston, MA, USA 109 Department of Plant Pathology, Irrigated Agricultural Research and Extension Center, Washington State University, Prosser, WA, USA 110 Utsunomiya University, Utsunomiya, Japan 111 Instituto de Biología Molecular y Celular de Plantas, Universitat Politècnica de València-Consejo Superior de Investigaciones Científicas, Valencia, Spain 112 School of Agriculture, Utsunomiya University, Utsunomiya, Japan 113 Novosibirsk State University, Novosibirsk, Novosibirsk Oblast, Russia 114 Department of Pathobiological Sciences, Influenza Research Institute, University of Wisconsin-Madison, Madison, USA 115 Institute of Virology, University of Veterinary Medicine Vienna, Vienna, Austria 116 College of Medicine, Mohammed Bin Rashid University of Medicine and Health Sciences, Dubai, United Arab Emirates 117 Institute for Biosecurity and Microbial Forensics. Stillwater, Oklahoma State University, Oklahoma, USA 118 Department of Microbiology, Icahn School of Medicine at Mount Sinai, New York, NY, USA 119 French Agency for Food, Environmental and Occupational Heath Safety ANSES, Laboratory of Ploufragan-Plouzané-Niort, Ploufragan, France 120 Instituto de Biología Molecular y Celular de Plantas (IBMCP), Consejo Superior de Investigaciones Cientificas-Universidat Politècnica de Valencia, Valencia, Spain 121 National Reference Centre for Arboviruses and Haemorrhagic Fever viruses, Department of Microbiology, Medical School, Aristotle University of Thessaloniki, Thessaloniki, Greece 122 Methods Development and Research Infrastructure, Bioinformatics and Systems Biology, Robert Koch Institute, Berlin, Germany 123 College of Veterinary Medicine, Baker Institute for Animal Health, Cornell University, Ithaca, NY, USA 124 Departamento de Veterinária, Universidade Federal de Viçosa, Viçosa, MG, Brazil 125 Center for Emerging Zoonotic and Parasitic Diseases, National Institute for Communicable Diseases of the National 2906 J. H. Kuhn et al. 1 3 Health Laboratory Service, Sandringham-Johannesburg, Gauteng, South Africa 126 Department of Population Health, College of Veterinary Medicine, University of Georgia, Athens, GA, USA 127 Institute of Diagnostic Virology, Friedrich-Loeffler-Institut, Greifswald-Insel Riems, Germany 128 Department of Microbiology and Immunology, Vagelos College of Physicians and Surgeons, Columbia University Irving Medical Center, New York, NY, USA 129 United States Army Medical Research Institute of Infectious Diseases (USAMRIID), Frederick, MD, USA 130 Instituto Biológico de São Paulo, São Paulo, SP, Brazil 131 Departamento de Biologia Celular, Universidade de Brasília, Brasília, Brazil 132 Instituto de Biotecnología y Biología Molecular, CONICET-UNLP, Facultad de Ciencias Exactas, Unversidad Nacional de La Plata, Buenos Aires, Argentina 133 Instituto de Biotecnología y Biología Molecular, CONICET-UNLP, Facultad de Ciencias Exactas, Unversidad Nacional de La Plata, Buenos Aires, Argentina 134 Institute of Diagnostic Virology, Friedrich-Loeffler-Institut, Greifswald-Insel Riems, Germany 135 Department of Infectious Disease & Global Health, Tufts University Cummings School of Veterinary Medicine, North Grafton, MA, USA 136 Institute for Fish and Wildlife Health, Vetsuisse Faculty, University of Bern, Bern, Switzerland 137 Department of Biochemistry, Molecular Biology, Entomology and Plant Pathology, Mississippi State University, Mississippi State, MS, USA 138 Institute for Plant Protection, National Agriculture and Food Research Organization, Tsukuba, Japan 139 Institute for Fish and Wildlife Health, Vetsuisse Faculty, University of Bern, Bern, Switzerland 140 Faculty of Medicine, University Medical Center-University Freiburg, Freiburg, Germany 141 Division of Neurological Sciences, Vetsuisse Faculty, University of Bern, Bern, Switzerland 142 Hawkesbury Institute for the Environment, Western Sydney University, Sydney, NSW, Australia 143 Sun Yat-sen University, Shenzhen, China 144 Bioinformatics Unit, Scientific Institute IRCCS “E. Medea”, Bosisio Parini, Italy 145 CBR Division, Dstl, Porton Down, Salisbury, Wiltshire, UK 146 Department of Microbiology, College of Medicine, Korea University, Seoul, Republic of Korea 147 School of Biomedical Sciences, Faculty of Health, Queensland University of Technology, Brisbane, QLD, Australia 148 Viral Special Pathogens Branch, Division of High-Consequence Pathogens and Pathology, Centers for Disease Control and Prevention, Atlanta, GA, USA 149 Department of Microbiology, Immunology, and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, CO, USA 150 Division of Global Epidemiology, International Institute for Zoonosis Control, Hokkaido University, Sapporo, Japan 151 Department of Botany and Zoology, Faculty of Science, Masaryk University, Brno, Czech Republic 152 Centers for Disease Control and Prevention, Atlanta, GA, USA 153 Laboratorio de Virología Molecular, Centro Ciencia & Vida, Fundación Ciencia & Vida and Facultad de Medicina y Ciencia, Universidad San Sebastián, Santiago, Chile 154 Institute for Plant Protection, National Agriculture and Food Research Organization, Tsukuba, Japan 155 Institute for Life and Medical Sciences (LiMe), Kyoto University, Kyoto, Japan 156 Institut Pasteur de Guinée, BP 4416, Conakry, Guinea 157 Aichi Agricultural Research Center, Nagakute, Aichi, Japan 158 Institute for Sustainable Plant Protection, National Research Council of Italy (IPSP-CNR), Torino, Italy 159 Department of Entomology and Plant Pathology, Division of Agriculture, University of Arkansas System, Fayetteville, AR, USA 160 Department of Viroscience, Erasmus MC, University Medical Centre Rotterdam, Rotterdam, The Netherlands 161 Department of Microbiology, Immunology and Transplantation, KU Leuven, Leuven, Belgium 162 Wageningen University and Research, Biointeractions and Plant Health, Wageningen, The Netherlands 163 Integrated Research Facility at Fort Detrick, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Frederick, MD, USA 164 National Biodefense Analysis and Countermeasures Center, Fort Detrick, Frederick, MD, USA 165 School of Chemistry and Molecular Biosciences, University of Queensland, St. Lucia, QLD, Australia 166 Department of Entomology and Plant Pathology, North Carolina State University, Raleigh, NC, USA 167 School of Medicine, University of Pittsburgh, Pittsburgh, PA, USA 168 National Center for Biotechnology Information, National Library of Medicine, National Institutes of Health, Bethesda, MD, USA 169 Environmental Agriculture Promotion Division, Department of Agricultural Development, Kochi Prefectural Government, Kochi, Kochi, Japan 170 Yokohama Plant Protection Station, Yokohama, Kanagawa, Japan 171 Institute of Insect Sciences, Zhejiang University, Hangzhou, China 172 National Institute for Communicable Disease Control and Prevention, China Center for Disease Control and Prevention, Beijing, China 173 Pharmaq Analytiq, Bergen, Norway